Chersobius

Genus of turtles

Chersobius

Speckled Cape tortoise (Chersobius signatus)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Superfamily:	Testudinoidea
Family:	Testudinidae
Genus:	Chersobius Fitzinger, 1835

Chersobius is a genus of tiny tortoises in the family Testudinidae, endemic to southern Africa. The genus includes the smallest tortoises in the world. All three species were previously assigned to the genus Homopus.

Naming

As a group, these closely related species are commonly known in Europe and Africa as padlopers (originally meaning "path-walkers" in Afrikaans), due to their habit of making tiny pathways through vegetation.^[1]^[2] In other parts of the world, such as the United States, they are known as Cape tortoises.

Distribution

The genus is indigenous and endemic to southern Africa, one within South Africa, one only in Namibia, and one possibly spanning across the border region of both countries.

Species

The genus contains these species:

Common Name	Scientific name	Distribution
Karoo padloper or Boulenger's cape tortoise	C. boulengeri	Karoo region^[3]
speckled padloper or speckled tortoise	C. signatus	South African west coast region, the smallest tortoise species in the world
Nama padloper or Berger's cape tortoise, previously H. bergeri	C. solus	southern Namibia^[4]

Conservation and captivity

They are threatened by habitat destruction, traffic on roads, overgrazing, and poaching for the pet trade.^[3]

Among the Chersobius species, C. signatus adapts well to captivity, as their diets are not highly specialized.^[2] The others do not generally survive well in captivity unless some effort is made to supply them with their natural food, that is, endemic plants from the Cape/Karoo regions.^[2] Many are taken from their natural habitat each year, and subsequently die as a result, as they do not readily adapt to typical captive diets and environment change.^[2] However, they can be very hardy in captivity, and most problems with captive care are caused by faulty nutrition, high humidity, or bad husbandry.^[2]

References

^ "Homopus Research Foundation, reports". Home.caiway.nl. Retrieved 2017-07-17.
^ ^a ^b ^c ^d ^e Corton, M., Homopus (Padloper Tortoise) Care, World Chelonian Trust (retrieved August 20, 2013).
^ ^a ^b "Homopus Research Foundation". Home.caiway.nl. Retrieved 2017-07-17.
^ "Nama Padloper Tortoise | Namibia". Namibian.org. Retrieved 2017-07-17.

Tortoise family

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines
Suborder: Cryptodira
Superfamily: Testudinoidea
Family: Testudinidae

Genera

Species of the tortoise family

Agrionemys

Russian tortoise

Chaco tortoise
Red-footed tortoise
Yellow-footed tortoise
†Chelonoidis alburyorum
†Chelonoidis cubensis
†Chelonoidis lutzae
†Chelonoidis monensis
†Chelonoidis sellovii
†Chelonoidis sombrerensis

Galápagos tortoise

Chelonoidis complex

†Cheirogaster

†Cheirogaster bacharidisi
†Cheirogaster gymnesica
†Cheirogaster schafferi

Chersina

Angulate tortoise

Chersobius

†Cylindraspis

Geochelone

Gopherus

†Hadrianus

†Hadrianus corsoni
†Hadrianus majusculus
†Hadrianus octonaria
†Hadrianus robustus
†Hadrianus schucherti
†Hadrianus utahensis
†Hadrianus vialovi

†Hesperotestudo

†Hesperotestudo percrassa

Pancake tortoise

Pyxis

Leopard tortoise

†Stylemys

†Stylemys botti
†Stylemys calaverensis
†Stylemys canetotiana
†Stylemys capax
†Stylemys conspecta
†Stylemys copei
†Stylemys emiliae
†Stylemys frizaciana
†Stylemys karakolensis
†Stylemys nebrascensis
†Stylemys neglectus
†Stylemys oregonensis
†Stylemys pygmea
†Stylemys uintensis
†Stylemys undabuna

Testudo

Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. Key: †=extinct.

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Suborder

Superfamily

Family

Genus

Cryptodira

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.