Eulipotyphla

Order of mammals

Eulipotyphla
Temporal range: Paleocene–Recent
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N
Clockwise from upper left: a solenodon, hedgehog,[a] mole and shrew
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Magnorder: Boreoeutheria
Superorder: Laurasiatheria
Order: Eulipotyphla
Waddell et al., 1999
Families
Synonyms

Eulipotyphla (/ˌjuːlɪpoʊˈtɪflə/, which means "truly fat and blind"[1]) is an order of mammals suggested by molecular methods of phylogenetic reconstruction, which includes the laurasiatherian members of the now-invalid polyphyletic order Lipotyphla, but not the afrotherian members (tenrecs, golden moles, and otter shrews, now in their own order Afrosoricida).

Eulipotyphla comprises the hedgehogs and gymnures (family Erinaceidae, formerly also the order Erinaceomorpha), solenodons (family Solenodontidae), the desmans, moles, and shrew-like moles (family Talpidae) and true shrews (family Soricidae). True shrews, talpids and solenodons were formerly grouped in Soricomorpha; however, Soricomorpha has been found to be paraphyletic, since erinaceids are the sister group of shrews.[2][3][4]

A 2023 study suggested that the order began to diversify prior to the K-Pg extinction, based on molecular clock estimates.[5]

Classification

European mole (Talpidae)
European hedgehog (Erinaceidae)
Common shrew (Soricidae)
Hispaniolan solenodon (Solenodontidae); solenodons are estimated to have diverged from other extant eulipotyphlans in the Late Cretaceous.[6][7]

Family-level cladogram of modern eulipotyphlan relationships, following Roca et al. and Brace et al.:[3][11]

The upper and lower basal subclades within the tree are the suborders Solenodonota and Erinaceota, respectively.[11] These two branches are estimated to have split ~72-74 million years (Ma) ago.[11][6][7] The Nesophontidae and Solenodontidae are thought to have separated roughly 57 Ma ago.[11] Split times for talpids vs. soricids plus erinaceids, and for soricids vs. erinaceids, have been estimated at 69 Ma and 64 Ma ago, respectively.[12]

Notes

References

  1. ^ Hassan, Mo (2009-10-11). "British Wildlife: N". The Disillusioned Taxonomist blog. Retrieved 2015-11-26.
  2. ^ Douady, C. J.; Chatelier, P. I.; Madsen, O.; de Jong, W. W.; Catzeflis, F.; Springer, M. S.; Stanhope, M. J. (October 2002). "Molecular phylogenetic evidence confirming the Eulipotyphla concept and in support of hedgehogs as the sister group to shrews". Molecular Phylogenetics and Evolution. 25 (1): 200–209. doi:10.1016/S1055-7903(02)00232-4. PMID 12383761.
  3. ^ a b Roca, A. L.; Bar-Gal, G. K.; Eizirik, E.; Helgen, K. M.; Maria, R.; Springer, M. S.; O'Brien, S. J.; Murphy, W. J. (2004-06-10). "Mesozoic origin for West Indian insectivores". Nature. 429 (6992): 649–651. Bibcode:2004Natur.429..649R. doi:10.1038/nature02597. PMID 15190349. S2CID 915633.
  4. ^ Bininda-Emonds, O. R. P.; Cardillo, M.; Jones, K. E.; MacPhee, R. D. E.; Beck, R. M. D.; Grenyer, R.; Price, S. A.; Vos, R. A.; Gittleman, J. L.; Purvis, A. (2007-03-29). "The delayed rise of present-day mammals". Nature. 446 (7135): 507–512. Bibcode:2007Natur.446..507B. doi:10.1038/nature05634. PMID 17392779. S2CID 4314965.
  5. ^ Foley, Nicole M.; Mason, Victor C.; Harris, Andrew J.; Bredemeyer, Kevin R.; Damas, Joana; Lewin, Harris A.; Eizirik, Eduardo; Gatesy, John; Karlsson, Elinor K.; Lindblad-Toh, Kerstin; Zoonomia Consortium‡; Springer, Mark S.; Murphy, William J.; Andrews, Gregory; Armstrong, Joel C. (2023-04-28). "A genomic timescale for placental mammal evolution". Science. 380 (6643): eabl8189. doi:10.1126/science.abl8189. ISSN 0036-8075. PMC 10233747. PMID 37104581.
  6. ^ a b de Lazaro, Enrico (19 March 2018). "Solenodon Genome Sequenced". Sci-News.com. Retrieved 2019-06-23.
  7. ^ a b Grigorev, K.; Kliver, S.; Dobrynin, P.; Komissarov, A.; Wolfsberger, W.; Krasheninnikova, K.; Afanador-Hernández, Y. M.; Brandt, A. L.; Paulino, L. A.; Carreras, R.; Rodríguez, L. E.; Núñez, A.; Brandt, J. R.; Silva, F.; Hernández-Martich, J. D.; Majeske, A. J.; Antunes, A.; Roca, A. L.; O'Brien, S. J.; Martínez-Cruzado, J. C.; Oleksyk, T. K. (2018). "Innovative assembly strategy contributes to understanding the evolution and conservation genetics of the endangered Solenodon paradoxus from the island of Hispaniola". GigaScience. 7 (6): giy025. doi:10.1093/gigascience/giy025. PMC 6009670. PMID 29718205.
  8. ^ Kim, N.H.; Lim, S.J.; Chae, H.M.; Park, Y.C. (2017). "Complete mitochondrial genome of the Amur hedgehog Erinaceus amurensis (Erinaceidae) and higher phylogeny of the family Erinaceidae". Genetics and Molecular Research. 16 (1). doi:10.4238/gmr16019300. PMID 28198504.
  9. ^ Dubey, S.; Salamin, N.; Ohdachi, S.D.; Barrière, P.; Vogel, P. (2007). "Molecular phylogenetics of shrews (Mammalia: Soricidae) reveal timing of transcontinental colonizations". Molecular Phylogenetics and Evolution. 44 (1): 126–137. doi:10.1016/j.ympev.2006.12.002. PMID 17267241. S2CID 3136125.
  10. ^ He, K.; Shinohara, A.; Helgen, K.M.; Springer, M.S.; Jiang, X.-L.; Campbell, K.L. (2017). "Talpid Mole Phylogeny Unites Shrew Moles and Illuminates Overlooked Cryptic Species Diversity". Molecular Biology and Evolution. 34 (1): 78–87. doi:10.1093/molbev/msw221. PMID 27795230.
  11. ^ a b c d Brace, S.; Thomas, J. A.; Dalén, L.; Burger, J.; MacPhee, R. D. E.; Barnes, I.; Turvey, S. T. (2016). "Evolutionary History of the Nesophontidae, the Last Unplaced Recent Mammal Family". Molecular Biology and Evolution. 33 (12): 3095–3103. doi:10.1093/molbev/msw186. PMID 27624716.
  12. ^ Springer, M. S.; Murphy, W. J.; Roca, A. L. (2018). "Appropriate fossil calibrations and tree constraints uphold the Mesozoic divergence of solenodons from other extant mammals". Molecular Phylogenetics and Evolution. 121: 158–165. doi:10.1016/j.ympev.2018.01.007. PMID 29331683.
  • v
  • t
  • e
Extant species of Eulipotyphla
Atelerix
  • Four-toed hedgehog (A. albiventris)
  • North African hedgehog (A. algirus)
  • Southern African hedgehog (A. frontalis)
  • Somali hedgehog (A. sclateri)
Erinaceus
  • Amur hedgehog (E. amurensis)
  • Southern white-breasted hedgehog (E. concolor)
  • West European hedgehog (E. europaeus)
  • Northern white-breasted hedgehog (E. roumanicus)
Hemiechinus
  • Long-eared hedgehog (H. auritus)
  • Indian long-eared hedgehog (H. collaris)
Mesechinus
  • Daurian hedgehog (M. dauuricus)
  • Hugh's hedgehog (M. hughi)
  • Gaoligong forest hedgehog (M. wangi)
  • Small-toothed forest hedgehog (M. miodon)
  • Eastern forest hedgehog (M. orientalis)
Paraechinus
  • Desert hedgehog (P. aethiopicus)
  • Brandt's hedgehog (P. hypomelas)
  • Indian hedgehog (P. micropus)
  • Bare-bellied hedgehog (P. nudiventris)
Echinosorex
  • Moonrat (E. gymnura)
Hylomys
  • Dwarf gymnure (H. parvus)
  • Javan short-tailed gymnure (H. suillus)
  • Bornean short-tailed gymnure (H. dorsalis)
  • Max's short-tailed gymnure (H. maxi)
  • Dalat gymnure (H. macarong)
  • Northern short-tailed gymnure (H. peguensis)
  • Leuser gymnure (H. vorax)
Neohylomys
  • Hainan gymnure (N. hainanensis)
Neotetracus
  • Shrew gymnure (N. sinensis)
Otohylomys
  • Long-eared gymnure (O. megalotis)
Podogymnura
  • Dinagat gymnure (P. aureospinula)
  • Mindanao gymnure (P. truei)
Crocidura
(White-toothed
shrews)
  • Cyrenaica shrew (C. aleksandrisi)
  • East African highland shrew (C. allex)
  • Andaman shrew (C. andamanensis)
  • C. annamitensis
  • Ansell's shrew (C. ansellorum)
  • Arabian shrew (C. arabica)
  • Jackass shrew (C. arispa)
  • Armenian shrew (C. armenica)
  • Asian gray shrew (C. attenuata)
  • Hun shrew (C. attila)
  • Bailey's shrew (C. baileyi)
  • Kinabalu shrew (C. baluensis)
  • Batak shrew (C. batakorum)
  • Mindanao shrew (C. beatus)
  • Beccari's shrew (C. beccarii)
  • Bottego's shrew (C. bottegi)
  • Bale shrew (C. bottegoides)
  • Thick-tailed shrew (C. brunnea)
  • Buettikofer's shrew (C. buettikoferi)
  • African dusky shrew (C. caliginea)
  • Canarian shrew (C. canariensis)
  • Caspian shrew (C. caspica)
  • Cinderella shrew (C. cinderella)
  • Congo white-toothed shrew (C. congobelgica)
  • C. cranbrooki
  • Long-footed shrew (C. crenata)
  • Crosse's shrew (C. crossei)
  • Reddish-gray musk shrew (C. cyanea)
  • Dent's shrew (C. denti)
  • Desperate shrew (C. desperata)
  • Dhofar shrew (C. dhofarensis)
  • Long-tailed musk shrew (C. dolichura)
  • Doucet's musk shrew (C. douceti)
  • Dsinezumi shrew (C. dsinezumi)
  • Eisentraut's shrew (C. eisentrauti)
  • Elgon shrew (C. elgonius)
  • Elongated shrew (C. elongata)
  • Heather shrew (C. erica)
  • Fischer's shrew (C. fischeri)
  • Greater red musk shrew (C. flavescens)
  • Flower's shrew (C. floweri)
  • Bornean shrew (C. foetida)
  • Fox's shrew (C. foxi)
  • Southeast Asian shrew (C. fuliginosa)
  • Savanna shrew (C. fulvastra)
  • Smoky white-toothed shrew (C. fumosa)
  • Bicolored musk shrew (C. fuscomurina)
  • Glass's shrew (C. glassi)
  • Gmelin's white-toothed shrew (C. gmelini)
  • Goliath shrew (C. goliath)
  • Peters's musk shrew (C. gracilipes)
  • Large-headed shrew (C. grandiceps)
  • Greater Mindanao shrew (C. grandis)
  • Grasse's shrew (C. grassei)
  • Luzon shrew (C. grayi)
  • Greenwood's shrew (C. greenwoodi)
  • C. guy
  • Harenna shrew (C. harenna)
  • C. hikmiya
  • Hildegarde's shrew (C. hildegardeae)
  • Hill's shrew (C. hilliana)
  • Lesser red musk shrew (C. hirta)
  • Andaman spiny shrew (C. hispida)
  • Horsfield's shrew (C. horsfieldii)
  • Hutan shrew (C. hutanis)
  • North African white-toothed shrew (C. ichnusae)
  • Indochinese shrew (C. indochinensis)
  • Jackson's shrew (C. jacksoni)
  • Jenkins's shrew (C. jenkinsi)
  • Jouvenet's shrew (C. jouvenetae)
  • Katinka's shrew (C. katinka)
  • Ke Go shrew (C. kegoensis)
  • Kivu shrew (C. kivuana)
  • Lamotte's shrew (C. lamottei)
  • Kivu long-haired shrew (C. lanosa)
  • Ussuri white-toothed shrew (C. lasiura)
  • Latona's shrew (C. latona)
  • Sulawesi shrew (C. lea)
  • Sumatran giant shrew (C. lepidura)
  • Bicolored shrew (C. leucodon)
  • Sulawesi tiny shrew (C. levicula)
  • Butiaba naked-tailed shrew (C. littoralis)
  • Savanna swamp shrew (C. longipes)
  • Lucina's shrew (C. lucina)
  • Ludia's shrew (C. ludia)
  • Moonshine shrew (C. luna)
  • Mauritanian shrew (C. lusitania)
  • MacArthur's shrew (C. macarthuri)
  • MacMillan's shrew (C. macmillani)
  • Nyiro shrew (C. macowi)
  • Malayan shrew (C. malayana)
  • Manenguba shrew (C. manengubae)
  • Makwassie musk shrew (C. maquassiensis)
  • Swamp musk shrew (C. mariquensis)
  • Gracile naked-tailed shrew (C. maurisca)
  • Javanese shrew (C. maxi)
  • Mindoro shrew (C. mindorus)
  • Sri Lankan long-tailed shrew (C. miya)
  • Kilimanjaro shrew (C. monax)
  • Sunda shrew (C. monticola)
  • Montane white-toothed shrew (C. montis)
  • West African long-tailed shrew (C. muricauda)
  • Mossy forest shrew (C. musseri)
  • Ugandan musk shrew (C. mutesae)
  • Somali dwarf shrew (C. nana)
  • Savanna dwarf shrew (C. nanilla)
  • Peninsular shrew (C. negligens)
  • Negros shrew (C. negrina)
  • Nicobar shrew (C. nicobarica)
  • Nigerian shrew (C. nigeriae)
  • Blackish white-toothed shrew (C. nigricans)
  • Black-footed shrew (C. nigripes)
  • African black shrew (C. nigrofusca)
  • Nimba shrew (C. nimbae)
  • Niobe's shrew (C. niobe)
  • West African pygmy shrew (C. obscurior)
  • African giant shrew (C. olivieri)
  • Oriental shrew (C. orientalis)
  • Ryukyu shrew (C. orii)
  • Palawan shrew (C. palawanensis)
  • Panay shrew (C. panayensis)
  • Sumatran long-tailed shrew (C. paradoxura)
  • Small-footed shrew (C. parvipes)
  • Sahelian tiny shrew (C. pasha)
  • Pale gray shrew (C. pergrisea)
  • Guramba shrew (C. phaeura)
  • C. phanluongi
  • Phu Hoc shrew (C. phuquocensis)
  • Cameroonian shrew (C. picea)
  • Pitman's shrew (C. pitmani)
  • Flat-headed shrew (C. planiceps)
  • Fraser's musk shrew (C. poensis)
  • Polia's shrew (C. polia)
  • Kashmir white-toothed shrew (C. pullata)
  • Rainey's shrew (C. raineyi)
  • Negev shrew (C. ramona)
  • Chinese white-toothed shrew (C. rapax)
  • Egyptian pygmy shrew (C. religiosa)
  • Sulawesi white-handed shrew (C. rhoditis)
  • Roosevelt's shrew (C. roosevelti)
  • Greater white-toothed shrew (C. russula)
  • Ugandan lowland shrew (C. selina)
  • Lesser rock shrew (C. serezkyensis)
  • Asian lesser white-toothed shrew (C. shantungensis)
  • Siberian shrew (C. sibirica)
  • Sicilian shrew (C. sicula)
  • Lesser gray-brown musk shrew (C. silacea)
  • Desert musk shrew (C. smithii)
  • Sokolov's shrew (C. sokolovi)
  • Somali shrew (C. somalica)
  • Kahuzi swamp shrew (C. stenocephala)
  • Lesser white-toothed shrew (C. suaveolens)
  • Iranian shrew (C. susiana)
  • Tanzanian shrew (C. tansaniana)
  • Tarella shrew (C. tarella)
  • Saharan shrew (C. tarfayensis)
  • Telford's shrew (C. telfordi)
  • Timor shrew (C. tenuis)
  • Thalia's shrew (C. thalia)
  • Therese's shrew (C. theresae)
  • São Tomé shrew (C. thomensis)
  • Christmas Island shrew (C. trichura)
  • Turbo shrew (C. turba)
  • Ultimate shrew (C. ultima)
  • Usambara shrew (C. usambarae)
  • Savanna path shrew (C. viaria)
  • Mamfe shrew (C. virgata)
  • Voi shrew (C. voi)
  • Voracious shrew (C. vorax)
  • Banka shrew (C. vosmaeri)
  • Lesser Ryukyu shrew (C. watasei)
  • Whitaker's shrew (C. whitakeri)
  • Wimmer's shrew (C. wimmeri)
  • Hainan Island shrew (C. wuchihensis)
  • Xanthippe's shrew (C. xantippe)
  • Yankari shrew (C. yankariensis)
  • Mikhail Zaitsev's shrew (C. zaitsevi )
  • Zaphir's shrew (C. zaphiri)
  • Zarudny's rock shrew (C. zarudnyi)
  • Upemba shrew (C. zimmeri)
  • Cretan shrew (C. zimmermanni)
Diplomesodon
  • Piebald shrew (D. pulchellus)
Feroculus
  • Kelaart's long-clawed shrew (F. feroculus)
Palawanosorex
  • Palawan moss shrew (P. muscorum)
Paracrocidura
(Large-headed
shrews)
  • Grauer's large-headed shrew (P. graueri)
  • Greater large-headed shrew (P. maxima)
  • Lesser large-headed shrew (P. schoutedeni)
Ruwenzorisorex
  • Ruwenzori shrew (R. suncoides)
Scutisorex
  • Hero shrew (S. somereni)
  • Thor's hero shrew (S. thori)
Solisorex
  • Pearson's long-clawed shrew (S. pearsoni)
Suncus
  • Taita shrew (S. aequatorius)
  • Black shrew (S. ater)
  • Day's shrew (S. dayi)
  • Etruscan shrew (S. etruscus)
  • Sri Lankan shrew (S. fellowesgordoni)
  • Bornean pygmy shrew (S. hosei)
  • Least dwarf shrew (S. infinitesimus)
  • Greater dwarf shrew (S. lixus)
  • Madagascan pygmy shrew (S. madagascariensis)
  • Malayan pygmy shrew (S. malayanus)
  • Climbing shrew (S. megalura)
  • Flores shrew (S. mertensi)
  • Asian highland shrew (S. montanus)
  • Asian house shrew (S. murinus)
  • Remy's pygmy shrew (S. remyi)
  • Anderson's shrew (S. stoliczkanus)
  • Lesser dwarf shrew (S. varilla)
  • Jungle shrew (S. zeylanicus)
Sylvisorex
(Forest shrews)
  • Dudu Akaibe's pygmy shrew (S. akaibei)
  • Cameroonian forest shrew (S. cameruniensis)
  • Corbet's forest shrew (S. corbeti)
  • Grant's forest shrew (S. granti)
  • Howell's forest shrew (S. howelli)
  • Bioko forest shrew (S. isabellae)
  • Johnston's forest shrew (S. johnstoni)
  • Kongana shrew (S. konganensis)
  • Moon forest shrew (S. lunaris)
  • Mount Cameroon forest shrew (S. morio)
  • Greater forest shrew (S. ollula)
  • Lesser forest shrew (S. oriundus)
  • Rain forest shrew (S. pluvialis)
  • Volcano shrew (S. vulcanorum)
Anourosoricini
Anourosorex
(Asian mole shrews)
  • Assam mole shrew (A. assamensis)
  • Giant mole shrew (A. schmidi)
  • Chinese mole shrew (A. squamipes)
  • Taiwanese mole shrew (A. yamashinai)
Blarinellini
Blarinella
(Asiatic short-tailed
shrews)
  • Indochinese short-tailed shrew (B. griselda)
  • Asiatic short-tailed shrew (B. quadraticauda)
  • Burmese short-tailed shrew (B. wardi)
Blarinini
Blarina
(American short-tailed
shrews)
  • Northern short-tailed shrew (B. brevicauda)
  • Southern short-tailed shrew (B. carolinensis)
  • Elliot's short-tailed shrew (B. hylophaga)
  • Everglades short-tailed shrew (B. peninsulae)
  • Sherman's short-tailed shrew (B. shermani)
Cryptotis
(Small-eared
shrews)
C. mexicana group
Mexican small-eared shrew (C. mexicana)
Nelson's small-eared shrew (C. nelsoni)
Grizzled Mexican small-eared shrew (C. obscura)
Phillips' small-eared shrew (C. phillipsii)
C. goldmani set
Central Mexican broad-clawed shrew (C. alticola)
Goldman's broad-clawed shrew (C. goldmani)
Goodwin's broad-clawed shrew (C. goodwini)
Guatemalan broad-clawed shrew (C. griseoventris)
C. lacertosus
C. mam
Oaxacan broad-clawed shrew (C. peregrina)
C. nigrescens group
Eastern Cordillera small-footed shrew (C. brachyonyx)
Colombian small-eared shrew (C. colombiana)
Honduran small-eared shrew (C. hondurensis)
Yucatan small-eared shrew (C. mayensis)
Darién small-eared shrew (C. mera)
Merriam's small-eared shrew (C. merriami)
Blackish small-eared shrew (C. nigrescens)
C. thomasi group
Southern Colombian small-eared shrew (C. andinus)
Ecuadorian small-eared shrew (C. equatoris)
Rainer's small-eared shrew (C. huttereri)
Medellín small-eared shrew (C. medellinia)
Merida small-eared shrew (C. meridensis)
Wandering small-eared shrew (C. montivaga)
Peruvian small-eared shrew (C. peruviensis)
Scaly-footed small-eared shrew (C. squaipes)
Tamá small-eared shrew (C. tamensis)
Thomas's small-eared shrew (C. thomasi)
C. parva group
Central American least shrew (C. orophila)
North American least shrew (C. parva)
Tropical small-eared shrew (C. tropicalis)
Ungrouped / relict
Enders's small-eared shrew (C. endersi)
Talamancan small-eared shrew (C. gracilis)
Big Mexican small-eared shrew (C. magna)
Nectogalini
Chimarrogale
(Asiatic water
shrews)
  • Malayan water shrew (C. hantu)
  • Himalayan water shrew (C. himalayica)
  • Bornean water shrew (C. phaeura)
  • Japanese water shrew (C. platycephalus)
  • Chinese water shrew (C. styani)
  • Sumatran water shrew (C. sumatrana)
Chodsigoa
  • Van Sung's shrew (C. caovansunga)
  • De Winton's shrew (C. hypsibia)
  • Lamulate shrew (C. lamula)
  • Lowe's shrew (C. parca)
  • Pygmy brown-toothed shrew (C. parva)
  • Salenski's shrew (C. salenskii)
  • Smith's shrew (C. smithii)
  • Lesser Taiwanese shrew (C. sodalis)
Episoriculus
  • Hodgsons's brown-toothed shrew (E. caudatus)
  • Taiwanese brown-toothed shrew (E. fumidus)
  • Long-tailed brown-toothed shrew (E. leucops)
  • Long-tailed mountain shrew (E. macrurus)
Nectogale
  • Elegant water shrew (N. elegans)
Neomys
  • Mediterranean water shrew (N. anomalus)
  • Eurasian water shrew (N. fodiens)
  • Transcaucasian water shrew (N. teres)
Soriculus
  • Himalayan shrew (S. nigrescens)
Notiosoricini
Megasorex
  • Mexican shrew (M. gigas)
Notiosorex
  • Cockrum's gray shrew (N. cockrumi)
  • Crawford's gray shrew (N. crawfordi)
  • Large-eared gray shrew (N. evotis)
  • Villa's gray shrew (N. villai)
Sorex
(Long-tailed
shrews)
Subgenus
Otisorex
  • Long-tailed shrew (S. dispar)
  • Smoky shrew (S. fumeus)
  • American pygmy shrew (S. hoyi)
  • Large-toothed shrew (S. macrodon)
  • Carmen mountain shrew (S. milleri)
  • Dwarf shrew (S. nanus)
  • Mexican long-tailed shrew (S. oreopolus)
  • Orizaba long-tailed shrew (S. orizabae)
  • Ornate shrew (S. ornatus)
  • Inyo shrew (S. tenellus)
  • Verapaz shrew (S. veraepacis)
S. vagrans complex
Glacier Bay water shrew (S. alaskanus)
Baird's shrew (S. bairdii)
Marsh shrew (S. bendirii)
Montane shrew (S. monticolus)
New Mexico shrew (S. neomexicanus)
Pacific shrew (S. pacificus)
American water shrew (S. palustris)
Fog shrew (S. sonomae)
Vagrant shrew (S. vagrans)
S. cinereus group
Kamchatka shrew (S. camtschatica)
Cinereus shrew (S. cinereus)
Prairie shrew (S. haydeni)
Saint Lawrence Island shrew (S. jacksoni)
Paramushir shrew (S. leucogaster)
Southeastern shrew (S. longirostris)
Mount Lyell shrew (S. lyelli)
Portenko's shrew (S. portenkoi)
Preble's shrew (S. preblei)
Pribilof Island shrew (S. pribilofensis)
Olympic shrew (S. rohweri)
Barren ground shrew (S. ugyunak)
Subgenus
Sorex
  • Dneper common shrew (S. averini)
  • Lesser striped shrew (S. bedfordiae)
  • Greater stripe-backed shrew (S. cylindricauda)
  • Chinese highland shrew (S. excelsus)
  • Azumi shrew (S. hosonoi)
  • Chinese shrew (S. sinalis)
  • Alaska tiny shrew (S. yukonicus)
S. alpinus group
Alpine shrew (S. alpinus)
Ussuri shrew (S. mirabilis)
S. araneus group
Valais shrew (S. antinorii)
Common shrew (S. araneus)
Udine shrew (S. arunchi)
Crowned shrew (S. coronatus)
Siberian large-toothed shrew (S. daphaenodon)
Iberian shrew (S. granarius)
Caucasian shrew (S. satunini)
S. arcticus group
Arctic shrew (S. arcticus)
Maritime shrew (S. maritimensis)
S. tundrensis group
Tien Shan shrew (S. asper)
Gansu shrew (S. cansulus)
Tundra shrew (S. tundrensis)
S. minutus group
Buchara shrew (S. buchariensis)
Kozlov's shrew (S. kozlovi)
Caucasian pygmy shrew (S. volnuchini)
S. caecutiens group
Laxmann's shrew (S. caecutiens)
Taiga shrew (S. isodon)
Eurasian least shrew (S. minutissimus)
Eurasian pygmy shrew (S. minutus)
Flat-skulled shrew (S. roboratus)
Shinto shrew (S. shinto)
Long-clawed shrew (S. unguiculatus)
S. gracillimus group
Slender shrew (S. gracillimus)
S. raddei group
Radde's shrew (S. raddei)
S. samniticus group
Apennine shrew (S. samniticus)
incertae sedis
  • Arizona shrew (S. arizonae)
  • Zacatecas shrew (S. emarginatus)
  • Merriam's shrew (S. merriami)
  • Kashmir pygmy shrew (S. planiceps)
  • Saussure's shrew (S. saussurei)
  • Sclater's shrew (S. sclateri)
  • San Cristobal shrew (S. stizodon)
  • Tibetan shrew (S. thibetanus)
  • Trowbridge's shrew (S. trowbridgii)
  • Chestnut-bellied shrew (S. ventralis)
  • Veracruz shrew (S. veraecrucis)
Congosorex
(Congo shrews)
  • Phillips's shrew (C. phillipsorum)
  • Greater Congo shrew (C. polli)
  • Lesser Congo shrew (C. verheyeni)
Myosorex
(Mouse shrews)
  • Babault's mouse shrew (M. babaulti)
  • Montane mouse shrew (M. blarina)
  • Bururi forest shrew (M. bururiensis)
  • Dark-footed mouse shrew (M. cafer)
  • Eisentraut's mouse shrew (M. eisentrauti)
  • Geata mouse shrew (M. geata)
  • Nyika mouse shrew (M. gnoskei)
  • Kihaule's mouse shrew (M. kihaulei)
  • Long-tailed forest shrew (M. longicaudatus)
  • Oku mouse shrew (M. okuensis)
  • Rumpi mouse shrew (M. rumpii)
  • Schaller's mouse shrew (M. schalleri)
  • Sclater's mouse shrew (M. sclateri)
  • Thin mouse shrew (M. tenuis)
  • Forest shrew (M. varius)
  • Kilimanjaro mouse shrew (M. zinki)
Surdisorex
(African mole
shrews)
  • Aberdare mole shrew (S. norae)
  • Mount Kenya mole shrew (S. polulus)
Scalopinae
(New World moles
and relatives)
Condylura
  • Star-nosed mole (C. cristata)
Parascalops
  • Hairy-tailed mole (P. breweri)
Scalopus
  • Eastern mole (S. aquaticus)
Scapanulus
  • Gansu mole (S. oweni)
Scapanus
(Western North
American moles)
  • Broad-footed mole (S. latimanus)
  • Coast mole (S. orarius)
  • Townsend's mole (S. townsendii)
Talpinae
(Old World moles
and relatives)
Desmana
  • Russian desman (D. moschata)
Dymecodon
  • True's shrew mole (D. pilirostris)
Euroscaptor
  • Greater Chinese mole (E. grandis)
  • Kloss's mole (E. klossi)
  • Long-nosed mole (E. longirostris)
  • Himalayan mole (E. micrura)
  • Small-toothed mole (E. parvidens)
  • Vietnamese mole (E. subanura)
Galemys
  • Pyrenean desman (G. pyrenaicus)
Mogera
  • Echigo mole (M. etigo)
  • Insular mole (M. insularis)
  • Kano mole (M. kanoana)
  • Kobe mole (M. kobeae)
  • Small Japanese mole (M. imaizumii)
  • Large mole (M. robusta)
  • Sado mole (M. tokudae)
  • Japanese mole (M. wogura)
  • Senkaku mole (M. uchidai)
Neurotrichus
  • American shrew mole (N. gibbsii)
Oreoscaptor
  • Japanese mountain mole (O. mizura)
Parascaptor
  • White-tailed mole (P. leucura)
Scaptochirus
  • Short-faced mole (S. moschatus)
Scaptonyx
  • Long-tailed mole (S. fusicaudus)
Talpa
  • Altai mole (T. altaica)
  • Aquitanian mole (T. aquitania)
  • Blind mole (T. caeca)
  • Caucasian mole (T. caucasica)
  • European mole (T. europaea)
  • Père David's mole (T. davidiana)
  • Levant mole (T. levantis)
  • Martino's mole (T. martinorum)
  • Spanish mole (T. occidentalis)
  • Ognev's mole (T. ognevi)
  • Roman mole (T. romana)
  • Balkan mole (T. stankovici)
  • Talysch mole (T. talyschensis)
Urotrichus
  • Japanese shrew mole (U. talpoides)
Uropsilinae
(Chinese shrew-like
moles)
  • Equivalent-teeth shrew mole (U. aequodonenia)
  • Anderson's shrew mole (U. andersoni)
  • Black-backed shrew mole (U. atronates)
  • Dabie Mountains shrew mole (U. dabieshanensis)
  • Gracile shrew mole (U. gracilis)
  • Inquisitive shrew mole (U. investigator)
  • Snow Mountain shrew mole (U. nivatus)
  • Chinese shrew mole (U. soricipes)
Atopogale
  • Cuban solenodon (A. cubana)
Solenodon
  • Hispaniolan solenodon (S. paradoxus)
Taxon identifiers
Eulipotyphla
Authority control databases: National Edit this at Wikidata
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  • United States


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