Gilmoremys

Extinct genus of turtles

Gilmoremys Temporal range: Upper Cretaceous, Maastrichtian PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	Trionychidae
Subfamily:	†Plastomeninae
Genus:	†Gilmoremys Joyce & Lyson, 2011
Species
†G. lancensis (Gilmore, 1928) [originally Aspideretes lancensis] (type) †G. gettyspherensis Joyce, Lyson, and Sertich, 2018

Gilmoremys is an extinct genus of softshell turtle which lived during the late Cretaceous (Maastrichtian stage) of North Dakota, Montana and Wyoming, United States.

Remains

Gilmoremys is known from five skulls, a mandible and an incomplete postcranial skeleton. The holotype of G. lancensis, USNM 6727, consists of a nearly complete carapace and an isolated hyoplastral fragment, was first assigned to the species Aspideretes lancensis. Many additional specimens were later discovered including cranial remains, and the material was assigned to its own genus, Gilmoremys. It was found from the Hell Creek Formation and from the Lance Formation (only the holotype). It was first named by Walter G. Joyce and Tyler R. Lyson in 2011 and the type species is Gilmoremys lancensis. The generic name honors Dr. Charles W. Gilmore.^[1] A second species, G. gettyspherensis, is known from the late Campanian Fruitland Formation of New Mexico.^[2]

Morphology

Juveniles of this species have narrow skulls, a narrow processus trochlearis oticum, a deep and narrow median palatal groove, low accessory ridges, and a secondary palate fully formed by the maxilla, skeletally mature individuals have notably broad skulls, a broad processus trochlearis oticum, a shallow but broad median palatal groove, high accessory ridges, and a substantial contribution of the vomer to the secondary palate. Two large trionychid skulls and isolated shell pieces have been discovered in deposits of the Hell Creek Formation exposed in Carter County, Montana, southeastern Montana. The skulls are noticeably larger than those previously discovered and differ in their overall form by being significantly broader. Detailed analysis reveals that the new skulls and shell fragments correspond with changes in overall form representing an ontogenetic shift towards the adult morphology of this species. ^[3]

Phylogeny

Cladogram after Joyce & Lyson, 2011:^[1]

Gilmoremys

Plastomenus

Hutchemys

	H. rememdium

	H. arctochelys

	H. tetanetron

	H. sterea

References

^ ^a ^b Walter G. Joyce and Tyler R. Lyson (2011). "New Material of Gilmoremys lancensis nov. comb. (Testudines: Trionychidae) from the Hell Creek Formation and the Diagnosis of Plastomenid Turtles". Journal of Paleontology. 85 (3): 442–459. doi:10.1666/10-127.1. S2CID 129918006.
^ Walter G. Joyce; Tyler R. Lyson; Joseph J.W. Sertich (2018). "A new species of trionychid turtle from the Upper Cretaceous (Campanian) Fruitland Formation of New Mexico, USA". Journal of Paleontology. 92 (6): 1107–1114. doi:10.1017/jpa.2018.30. S2CID 134161813.
^ Joyce, Walter; Lyson, Tyler; Williams, Scott (2016). "New Cranial Material of Gilmoremys lancensis (Testudines, Trionychidae) from the Hell Creek Formation of Southeastern Montana, U.S.A.". Journal of Vertebrate Paleontology. 36 (6): e1225748. doi:10.1080/02724634.2016.1225748. S2CID 133358179.

Trionychidae family

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines
Superfamily: Trionychia

Genera

Species of the Trionychidae family

Amyda

Asiatic softshell turtle
†Amyda gregaria
Amyda ornata

Apalone

†Axestemys

†Axestemys byssina
†Axestemys cerevisia
†Axestemys montinsana
†Axestemys quinni
†Axestemys splendida
†Axestemys vittata

Chitra

Cyclanorbis

Cycloderma

Dogania

Malayan softshell turtle

†Gilmoremys

†Gilmoremys gettyspherensis
†Gilmoremys lancensis

†Hutchemys

†Hutchemys arctochelys
†Hutchemys rememdium
†Hutchemys tetanetron
†Hutchemys sterea
†Hutchemys acupictus

†Khunnuchelys

†Khunnuchelys erinhotensis
†Khunnuchelys kizylkumensis
†Khunnuchelys lophorhothon

Lissemys

Nilssonia

†Palaeoamyda

†Palaeoamyda messeliana

Palea

Wattle-necked softshell turtle

Pelochelys

Pelodiscus

Rafetus

Trionyx

Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. †=extinct.

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Suborder

Superfamily

Family

Genus

Cryptodira

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.