Goral

Genus of mammals

Goral[1]
Chinese goral, Nemorhaedus griseus
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Bovidae
Subfamily: Caprinae
Tribe: Ovibovini
Genus: Naemorhedus
Hamilton Smith, 1827
Type species
Antilope goral[1]
Hardwicke, 1825
Species

Naemorhedus goral
Naemorhedus caudatus
Naemorhedus baileyi
Naemorhedus griseus

The gorals are four species in the genus Naemorhedus. They are small ungulates with a goat-like or antelope-like appearance. Until recently, this genus also contained the serow species (now in genus Capricornis).[1]

Etymology

The original name is based on Latin nemor-haedus, from nemus, nemoris 'grove' and haedus 'little goat', but it was misspelt Naemorhedus by Hamilton Smith (1827).[2][3]

The name "goral" comes from an eastern Indian word for the Himalayan goral.

Extant species

Image Scientific name Common Name Distribution
Naemorhedus goral Himalayan goral (also known as ghural) northern Pakistan, northwest and northeast India, as well as Nepal, Bhutan, and southern Tibet
Naemorhedus caudatus long-tailed goral eastern Russia and northeast China through North Korea and South Korea. A population has also been documented in the Demilitarized Zone on the Korean Peninsula
Naemorhedus baileyi red goral Yunnan province of China, to Tibet and northeast India through northern Myanmar
Naemorhedus griseus Chinese goral China, northeast India, Myanmar, northwest Thailand, northern Vietnam, and possibly Laos.

Habitat

Gorals are often found on rocky hillsides at high elevations. Though their territories often coincide with those of the closely related serow, the goral will usually be found on higher, steeper slopes with less vegetation.

Characteristics

Gorals typically weigh 25–40 kilograms (55–88 lb) and are 80–130 centimetres (31–51 in) in length, with short, backward-facing horns. Coloration differs between species and individuals, but generally ranges from light gray to dark red-brown, with lighter patches on the chest, throat, and underside, and a dark stripe down the spine. They have woolly undercoats covered by longer, coarser hair, which helps to protect them in the cold areas where they are often found.

Though the groups share many similarities, gorals are stockier than antelopes and have broader, heavier hooves. Female gorals have four functional teats, while female goats and sheep have only two functional teats. Unlike serows, gorals have no working preorbital glands.

References

  1. ^ a b c Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN 0-8018-8221-4. OCLC 62265494.
  2. ^ Groves, C. P., and Grubb, P., 1985, Reclassification of the serows and gorals (Nemorhaedus: Bovidae). In The Biology and Management of Mountain Ungulates. Edited by S. Lovari. London: Croom Helm. pp. 45-50.
  3. ^ Article 32.5.1. Archived 2016-03-14 at the Wayback Machine of International Commission on Zoological Nomenclature reads: "If there is in the original publication itself, without recourse to any external source of information, clear evidence of an inadvertent error, such as a lapsus calami or a copyist's or printer's error, it must be corrected."
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Extant Artiodactyla species
Suborder Ruminantia
Antilocapridae
Antilocapra
  • Pronghorn (A. americana)
Giraffidae
Okapia
  • Okapi (O. johnstoni)
Giraffa
  • Northern giraffe (G. camelopardalis)
  • Southern giraffe (G. giraffa)
  • Reticulated giraffe (G. reticulata)
  • Masai giraffe (G. tippelskirchi)
Moschidae
Moschus
  • Anhui musk deer (M. anhuiensis)
  • Dwarf musk deer (M. berezovskii)
  • Alpine musk deer (M. chrysogaster)
  • Kashmir musk deer (M. cupreus)
  • Black musk deer (M. fuscus)
  • Himalayan musk deer (M. leucogaster)
  • Siberian musk deer (M. moschiferus)
Tragulidae
Hyemoschus
  • Water chevrotain (H. aquaticus)
Moschiola
  • Indian spotted chevrotain (M. indica)
  • Yellow-striped chevrotain (M. kathygre)
  • Sri Lankan spotted chevrotain (M. meminna)
Tragulus
  • Java mouse-deer (T. javanicus)
  • Lesser mouse-deer (T. kanchil)
  • Greater mouse-deer (T. napu)
  • Philippine mouse-deer (T. nigricans)
  • Vietnam mouse-deer (T. versicolor)
  • Williamson's mouse-deer (T. williamsoni)
Cervidae
Large family listed below
Bovidae
Large family listed below
Family Cervidae
Cervinae
Muntiacus
  • Bornean yellow muntjac (M. atherodes)
  • Hairy-fronted muntjac (M. crinifrons)
  • Fea's muntjac (M. feae)
  • Gongshan muntjac (M. gongshanensis)
  • Sumatran muntjac (M. montanus)
  • Southern red muntjac (M. muntjak)
  • Pu Hoat muntjac (M. puhoatensis)
  • Leaf muntjac (M. putaoensis)
  • Reeves's muntjac (M. reevesi)
  • Roosevelt's muntjac (M. rooseveltorum)
  • Truong Son muntjac (M. truongsonensis)
  • Northern red muntjac (M. vaginalis)
  • Giant muntjac (M. vuquangensis)
Elaphodus
  • Tufted deer (E. cephalophus)
Dama
  • European fallow deer (D. dama)
  • Persian fallow deer (D. mesopotamica)
Axis
  • Chital (A. axis)
  • Calamian deer (A. calamianensis)
  • Bawean deer (A. kuhlii)
  • Hog deer (A. porcinus)
Rucervus
  • Barasingha (R. duvaucelii)
  • Eld's deer (R. eldii)
Elaphurus
  • Père David's deer (E. davidianus)
Rusa
  • Visayan spotted deer (R. alfredi)
  • Philippine sambar (R. mariannus)
  • Rusa deer (R. timorensis)
  • Sambar (R. unicolor)
Cervus
  • Thorold's deer (C. albirostris)
  • Red deer (C. elaphus)
  • Elk (C. canadensis)
  • Central Asian red deer (C. hanglu)
  • Sika deer (C. nippon)
Capreolinae
Alces
  • Moose (A. alces)
Hydropotes
  • Water deer (H. inermis)
Capreolus
  • European roe deer (C. capreolus)
  • Siberian roe deer (C. pygargus)
Rangifer
  • Reindeer (R. tarandus)
Hippocamelus
  • Taruca (H. antisensis)
  • South Andean deer (H. bisulcus)
Mazama
  • Red brocket (M. americana)
  • Small red brocket (M. bororo)
  • Merida brocket (M. bricenii)
  • Dwarf brocket (M. chunyi)
  • Gray brocket (M. gouazoubira)
  • Pygmy brocket (M. nana)
  • Amazonian brown brocket (M. nemorivaga)
  • Little red brocket (M. rufina)
  • Central American red brocket (M. temama)
Ozotoceros
  • Pampas deer (O. bezoarticus)
Blastocerus
  • Marsh deer (B. dichotomus)
Pudu
  • Northern pudu (P. mephistophiles)?
  • Southern pudu (P. pudu)
Pudella?
  • Peruvian Yungas pudu (P. carlae)
  • Northern pudu (P. mephistophiles)
Odocoileus
  • Mule deer (O. hemionus)
  • Yucatan brown brocket (O. pandora)
  • White-tailed deer (O. virginianus)
Family Bovidae
Hippotraginae
Hippotragus
  • Roan antelope (H. equinus)
  • Sable antelope (H. niger)
Oryx
  • East African oryx (O. beisa)
  • Scimitar oryx (O. dammah)
  • Gemsbok (O. gazella)
  • Arabian oryx (O. leucoryx)
Addax
  • Addax (A. nasomaculatus)
Reduncinae
Kobus
  • Waterbuck (K. ellipsiprymnus)
  • Kob (K. kob)
  • Lechwe (K. leche)
  • Nile lechwe (K. megaceros)
  • Puku (K. vardonii)
Redunca
  • Southern reedbuck (R. arundinum)
  • Mountain reedbuck (R. fulvorufula)
  • Bohor reedbuck (R. redunca)
Aepycerotinae
Aepyceros
  • Impala (A. melampus)
Peleinae
Pelea
  • Grey rhebok (P. capreolus)
Alcelaphinae
Beatragus
  • Hirola (B. hunteri)
Damaliscus
  • Common tsessebe (D. lunatus)
  • Bontebok (D. pygargus)
Alcelaphus
  • Hartebeest (A. buselaphus)
Connochaetes
  • Black wildebeest (C. gnou)
  • Blue wildebeest (C. taurinus)
Pantholopinae
Pantholops
  • Tibetan antelope (P. hodgsonii)
Caprinae
Large subfamily listed below
Bovinae
Large subfamily listed below
Antilopinae
Large subfamily listed below
Family Bovidae (subfamily Caprinae)
Ammotragus
  • Barbary sheep (A. lervia)
Arabitragus
  • Arabian tahr (A. jayakari)
Budorcas
  • Takin (B. taxicolor)
Capra
  • Wild goat (C. aegagrus)
  • West Caucasian tur (C. caucasia)
  • East Caucasian tur (C. cylindricornis)
  • Markhor (C. falconeri)
  • Domestic goat (C. hircus)
  • Alpine ibex (C. ibex)
  • Nubian ibex (C. nubiana)
  • Iberian ibex (C. pyrenaica)
  • Siberian ibex (C. sibirica)
  • Walia ibex (C. walie)
Capricornis
  • Japanese serow (C. crispus)
  • Red serow (C. rubidus)
  • Mainland serow (C. sumatraensis)
  • Taiwan serow (C. swinhoei)
Hemitragus
  • Himalayan tahr (H. jemlahicus)
Naemorhedus
  • Red goral (N. baileyi)
  • Long-tailed goral (N. caudatus)
  • Himalayan goral (N. goral)
  • Chinese goral (N. griseus)
Oreamnos
  • Mountain goat (O. americanus)
Ovibos
  • Muskox (O. moschatus)
Nilgiritragus
  • Nilgiri tahr (N. hylocrius)
Ovis
  • Argali (O. ammon)
  • Domestic sheep (O. aries)
  • Bighorn sheep (O. canadensis)
  • Dall sheep (O. dalli)
  • Mouflon (O. gmelini)
  • Snow sheep (O. nivicola)
  • Urial (O. vignei)
Pseudois
  • Bharal (P. nayaur)
Rupicapra
  • Pyrenean chamois (R. pyrenaica)
  • Chamois (R. rupicapra)
Family Bovidae (subfamily Bovinae)
Boselaphini
Tetracerus
  • Four-horned antelope (T. quadricornis)
Boselaphus
  • Nilgai (B. tragocamelus)
Bovini
Bubalus
  • Wild water buffalo (B. arnee)
  • Domestic water buffalo (B. bubalis)
  • Lowland anoa (B. depressicornis)
  • Tamaraw (B. mindorensis)
  • Mountain anoa (B. quarlesi)
Bos
  • American bison (B. bison)
  • European bison (B. bonasus)
  • Bali cattle (B. domesticus)
  • Gayal (B. frontalis)
  • Gaur (B. gaurus)
  • Domestic yak (B. grunniens)
  • Zebu (B. indicus)
  • Banteng (B. javanicus)
  • Wild yak (B. mutus)
  • Cattle (B. taurus)
Pseudoryx
  • Saola (P. nghetinhensis)
Syncerus
  • African buffalo (S. caffer)
Tragelaphini
Tragelaphus
(including kudus)
  • Nyala (T. angasii)
  • Mountain nyala (T. buxtoni)
  • Bongo (T. eurycerus)
  • Lesser kudu (T. imberbis)
  • Harnessed bushbuck (T. scriptus)
  • Sitatunga (T. spekeii)
  • Greater kudu (T. strepsiceros)
  • Cape bushbuck (T. sylvaticus)
Taurotragus
  • Giant eland (T. derbianus)
  • Common eland (T. oryx)
Family Bovidae (subfamily Antilopinae)
Antilopini
Ammodorcas
  • Dibatag (A. clarkei)
Antidorcas
  • Springbok (A. marsupialis)
Antilope
  • Blackbuck (A. cervicapra)
Eudorcas
  • Mongalla gazelle (E. albonotata)
  • Red-fronted gazelle (E. rufifrons)
  • Thomson's gazelle (E. thomsonii)
  • Heuglin's gazelle (E. tilonura)
Gazella
  • Chinkara (G. bennettii)
  • Cuvier's gazelle (G. cuvieri)
  • Dorcas gazelle (G. dorcas)
  • Erlanger's gazelle (G. erlangeri)
  • Mountain gazelle (G. gazella)
  • Rhim gazelle (G. leptoceros)
  • Speke's gazelle (G. spekei)
  • Goitered gazelle (G. subgutturosa)
Litocranius
  • Gerenuk (L. walleri)
Nanger
  • Dama gazelle (N. dama)
  • Grant's gazelle (N. granti)
  • Bright's gazelle (N. notatus)
  • Peter's gazelle (N. petersii)
  • Soemmerring's gazelle (N. soemmerringii)
Procapra
  • Mongolian gazelle (P. gutturosa)
  • Goa (P. picticaudata)
  • Przewalski's gazelle (P. przewalskii)
Saigini
Pantholops
  • Tibetan antelope (P. hodgsonii)
Saiga
  • Saiga antelope (S. tatarica)
Neotragini
Dorcatragus
  • Beira (D. megalotis)
Madoqua
  • Günther's dik-dik (M. guentheri)
  • Kirk's dik-dik (M. kirkii)
  • Silver dik-dik (M. piacentinii)
  • Salt's dik-dik (M. saltiana)
Neotragus
  • Bates' pygmy antelope (N. batesi)
  • Suni (N. moschatus)
  • Royal antelope (N. pygmaeus)
Oreotragus
  • Klipspringer (O. oreotragus)
Ourebia
  • Oribi (O. ourebi)
Raphicerus
  • Steenbok (R. campestris)
  • Cape grysbok (R. melanotis)
  • Sharpe's grysbok (R. sharpei)
Cephalophini
Cephalophus
  • Aders's duiker (C. adersi)
  • Brooke's duiker (C. brookei)
  • Peters' duiker (C. callipygus)
  • White-legged duiker (C. crusalbum)
  • Bay duiker (C. dorsalis)
  • Harvey's duiker (C. harveyi)
  • Jentink's duiker (C. jentinki)
  • White-bellied duiker (C. leucogaster)
  • Red forest duiker (C. natalensis)
  • Black duiker (C. niger)
  • Black-fronted duiker (C. nigrifrons)
  • Ogilby's duiker (C. ogilbyi)
  • Ruwenzori duiker (C. rubidis)
  • Red-flanked duiker (C. rufilatus)
  • Yellow-backed duiker (C. silvicultor)
  • Abbott's duiker (C. spadix)
  • Weyns's duiker (C. weynsi)
  • Zebra duiker (C. zebra)
Philantomba
  • Blue duiker (P. monticola)
  • Maxwell's duiker (P. maxwellii)
  • Walter's duiker (P. walteri)
Sylvicapra
  • Common duiker (S. grimmia)
Suborder Suina
Suidae
Babyrousa
  • Buru babirusa (B. babyrussa)
  • North Sulawesi babirusa (B. celebensis)
  • Togian babirusa (B. togeanensis)
Hylochoerus
  • Giant forest hog (H. meinertzhageni)
Phacochoerus
  • Desert warthog (P. aethiopicus)
  • Common warthog (P. africanus)
Porcula
  • Pygmy hog (P. salvania)
Potamochoerus
  • Bushpig (P. larvatus)
  • Red river hog (P. porcus)
Sus
  • Palawan bearded pig (S. ahoenobarbus)
  • Bornean bearded pig (S. barbatus)
  • Visayan warty pig (S. cebifrons)
  • Celebes warty pig (S. celebensis)
  • Domestic pig (S. domesticus)
  • Flores warty pig (S. heureni)
  • Oliver's warty pig (S. oliveri)
  • Philippine warty pig (S. philippensis)
  • Wild boar (S. scrofa)
  • Timor warty pig (S. timoriensis)
  • Javan warty pig (S. verrucosus)
Tayassuidae
Tayassu
  • White-lipped peccary (T. pecari)
Catagonus
  • Chacoan peccary (C. wagneri)
Dicotyles
  • Collared peccary (D. tajacu)
Suborder Tylopoda
Camelidae
Lama
  • Llama (L. glama)
  • Guanaco (L. guanicoe)
  • Alpaca (L. pacos)
  • Vicuña (L. vicugna)
Camelus
  • Domestic Bactrian camel (C. bactrianus)
  • Dromedary/Arabian camel (C. dromedarius)
  • Wild Bactrian camel (C. ferus)
Suborder Whippomorpha
Hippopotamidae
Hippopotamus
  • Hippopotamus (H. amphibius)
Choeropsis
  • Pygmy hippopotamus (C. liberiensis)
Cetacea
  • see Cetacea
Taxon identifiers
Naemorhedus