Gymnure

Subfamily of mammals

Gymnures & moonrats[1]
Bornean short-tailed gymnure
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Eulipotyphla
Family: Erinaceidae
Subfamily: Galericinae
Pomel, 1848
Genera[5]
  • Apulogalerix[2]
  • Deinogalerix
  • Dzavui[3]
  • Echinosorex
  • Eochenus
  • Eogalericius
  • Galerix
  • Hylomys
  • Lantanotherium
  • Microgalericulus
  • Neohylomys
  • Neotetracus
  • Neurogymnurus
  • Ocajila
  • Oligochenus
  • Parasorex
  • Podogymnura
  • Proterix
  • Protogalericius
  • Pseudoneurogymnurus
  • Riddleria[4]
  • Schizogalerix
  • Tetracus
  • Thaiagymnura

Gymnures, also called hairy hedgehogs or moonrats, are mammals belonging to the subfamily Galericinae, in the family Erinaceidae and the order Eulipotyphla. Gymnures resemble rats but are not closely related as they are not rodents; they are instead closely related to hedgehogs, which also belong to Erinaceidae. They are thought to have appeared in Eastern Asia before their closest relatives, and changed little from the original ancestor, which is thought to have been also the ancestor of the shrews.

Description

Although the gymnures are more closely related to the hedgehogs, full-grown gymnures superficially resemble large rats, shrews, and opossums.

The gymnure's body plan is believed to resemble that of the earliest mammals,[citation needed] with a large, toothy head about 1/3 the length of the total body, a naked furless tail for balance and thermoregulatory purposes, and a plantigrade stance. In direct contrast to the closely related hedgehogs, gymnures are not spiny.[6]

They also have an outstanding sense of smell, and tactile response in the snout region.

Distribution and habitat

Gymnures inhabit moist jungle terrain in various locales of Southeast Asia, including Vietnam, Sumatra, China and the Malay Peninsula.

Behaviour

Gymnures are primarily carnivorous. They are nocturnal or crepuscular: they come out to forage at twilight or in the night to search the forest floor, using smell to find the animals that they eat. Gymnures eat various arthropods, mice, small reptiles and amphibians, with occasional fruit and fungi.

Gymnures keep territories, and individuals are solitary except when breeding. Gymnures have a very strong scent, typically described as a rancid garlic or onion smell, which is produced by its territory-marking scent glands. Several creatures similar in form and niche, such as the opossum and solenodon, have an odor similar to the gymnure's.

Classification

This subfamily has alternately been called Echinosoricinae, Galericinae, and Hylomyinae. Some researchers prefer Hylomyinae because the specific relationships of the extinct genus Galerix to living erinaceids are uncertain.[7] There are fifteen extant species in six genera:[1][8]

References

  1. ^ a b Hutterer, R. (2005). Wilson, D.E.; Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 212–217. ISBN 978-0-8018-8221-0. OCLC 62265494.
  2. ^ Masini, F.; Fanfani, F. (2013). "Apulogalerix pusillus nov. gen., nov. sp., the small-sized Galericinae (Erinaceidae, Mammalia) from the "Terre Rosse" fissure filling of the Gargano (Foggia, South-Eastern Italy)". Geobios. 46 (1–2): 89–104. doi:10.1016/j.geobios.2012.10.008.
  3. ^ Jiménez-Hidalgo, E.; Guerrero-Arenas, R.; Crespo, V. D. (2022). "First galericine erinaceid (Mammalia: Eulipotyphla) from the early Oligocene of tropical North America". Historical Biology: 1–6. doi:10.1080/08912963.2022.2070018. S2CID 248467861.
  4. ^ van den Hoek Ostende, L. W. (2003). "Riddleria atecensis nov. gen. nov. sp., a peculiar erinaceid (Erinaceomorpha, Mammalia) from the Lower Miocene of Spain". Beiträge zur Paläontologie. 28: 1–11.
  5. ^ Lopatin, A. V. (2006). "Early Paleogene insectivore mammals of Asia and establishment of the major groups of Insectivora". Paleontological Journal. 40 (S3): S205–S405. doi:10.1134/S0031030106090012. S2CID 84144565.
  6. ^ Feldhamer, George [and 4 others] (2015). Mammalogy: Adaptation, Diversity, Ecology, Edition 4. Baltimore: Johns Hopkins University Press. p. 285. ISBN 978-1-4214-1588-8.{{cite book}}: CS1 maint: numeric names: authors list (link)
  7. ^ Gould, G.C. (1995). "Hedgehog phylogeny (Mammalia, Erinaceidae) – the reciprocal illumination of the quick and the dead". American Museum Novitates (3131): 1–45. hdl:2246/3665.
  8. ^ Hinckley, A.; Camacho-Sanchez, M.; et al. (2023). "An integrative taxonomic revision of lesser gymnures (Eulipotyphla: Hylomys) reveals five new species and emerging patterns of local endemism in Tropical East Asia". Zoological Journal of the Linnean Society: zlad177. doi:10.1093/zoolinnean/zlad177.
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Extant species of Eulipotyphla
Atelerix
  • Four-toed hedgehog (A. albiventris)
  • North African hedgehog (A. algirus)
  • Southern African hedgehog (A. frontalis)
  • Somali hedgehog (A. sclateri)
Erinaceus
  • Amur hedgehog (E. amurensis)
  • Southern white-breasted hedgehog (E. concolor)
  • West European hedgehog (E. europaeus)
  • Northern white-breasted hedgehog (E. roumanicus)
Hemiechinus
  • Long-eared hedgehog (H. auritus)
  • Indian long-eared hedgehog (H. collaris)
Mesechinus
  • Daurian hedgehog (M. dauuricus)
  • Hugh's hedgehog (M. hughi)
  • Gaoligong forest hedgehog (M. wangi)
  • Small-toothed forest hedgehog (M. miodon)
  • Eastern forest hedgehog (M. orientalis)
Paraechinus
  • Desert hedgehog (P. aethiopicus)
  • Brandt's hedgehog (P. hypomelas)
  • Indian hedgehog (P. micropus)
  • Bare-bellied hedgehog (P. nudiventris)
Echinosorex
  • Moonrat (E. gymnura)
Hylomys
  • Dwarf gymnure (H. parvus)
  • Javan short-tailed gymnure (H. suillus)
  • Bornean short-tailed gymnure (H. dorsalis)
  • Max's short-tailed gymnure (H. maxi)
  • Dalat gymnure (H. macarong)
  • Northern short-tailed gymnure (H. peguensis)
  • Leuser gymnure (H. vorax)
Neohylomys
  • Hainan gymnure (N. hainanensis)
Neotetracus
  • Shrew gymnure (N. sinensis)
Otohylomys
  • Long-eared gymnure (O. megalotis)
Podogymnura
  • Dinagat gymnure (P. aureospinula)
  • Mindanao gymnure (P. truei)
Crocidura
(White-toothed
shrews)
  • Cyrenaica shrew (C. aleksandrisi)
  • East African highland shrew (C. allex)
  • Andaman shrew (C. andamanensis)
  • C. annamitensis
  • Ansell's shrew (C. ansellorum)
  • Arabian shrew (C. arabica)
  • Jackass shrew (C. arispa)
  • Armenian shrew (C. armenica)
  • Asian gray shrew (C. attenuata)
  • Hun shrew (C. attila)
  • Bailey's shrew (C. baileyi)
  • Kinabalu shrew (C. baluensis)
  • Batak shrew (C. batakorum)
  • Mindanao shrew (C. beatus)
  • Beccari's shrew (C. beccarii)
  • Bottego's shrew (C. bottegi)
  • Bale shrew (C. bottegoides)
  • Thick-tailed shrew (C. brunnea)
  • Buettikofer's shrew (C. buettikoferi)
  • African dusky shrew (C. caliginea)
  • Canarian shrew (C. canariensis)
  • Caspian shrew (C. caspica)
  • Cinderella shrew (C. cinderella)
  • Congo white-toothed shrew (C. congobelgica)
  • C. cranbrooki
  • Long-footed shrew (C. crenata)
  • Crosse's shrew (C. crossei)
  • Reddish-gray musk shrew (C. cyanea)
  • Dent's shrew (C. denti)
  • Desperate shrew (C. desperata)
  • Dhofar shrew (C. dhofarensis)
  • Long-tailed musk shrew (C. dolichura)
  • Doucet's musk shrew (C. douceti)
  • Dsinezumi shrew (C. dsinezumi)
  • Eisentraut's shrew (C. eisentrauti)
  • Elgon shrew (C. elgonius)
  • Elongated shrew (C. elongata)
  • Heather shrew (C. erica)
  • Fischer's shrew (C. fischeri)
  • Greater red musk shrew (C. flavescens)
  • Flower's shrew (C. floweri)
  • Bornean shrew (C. foetida)
  • Fox's shrew (C. foxi)
  • Southeast Asian shrew (C. fuliginosa)
  • Savanna shrew (C. fulvastra)
  • Smoky white-toothed shrew (C. fumosa)
  • Bicolored musk shrew (C. fuscomurina)
  • Glass's shrew (C. glassi)
  • Gmelin's white-toothed shrew (C. gmelini)
  • Goliath shrew (C. goliath)
  • Peters's musk shrew (C. gracilipes)
  • Large-headed shrew (C. grandiceps)
  • Greater Mindanao shrew (C. grandis)
  • Grasse's shrew (C. grassei)
  • Luzon shrew (C. grayi)
  • Greenwood's shrew (C. greenwoodi)
  • C. guy
  • Harenna shrew (C. harenna)
  • C. hikmiya
  • Hildegarde's shrew (C. hildegardeae)
  • Hill's shrew (C. hilliana)
  • Lesser red musk shrew (C. hirta)
  • Andaman spiny shrew (C. hispida)
  • Horsfield's shrew (C. horsfieldii)
  • Hutan shrew (C. hutanis)
  • North African white-toothed shrew (C. ichnusae)
  • Indochinese shrew (C. indochinensis)
  • Jackson's shrew (C. jacksoni)
  • Jenkins's shrew (C. jenkinsi)
  • Jouvenet's shrew (C. jouvenetae)
  • Katinka's shrew (C. katinka)
  • Ke Go shrew (C. kegoensis)
  • Kivu shrew (C. kivuana)
  • Lamotte's shrew (C. lamottei)
  • Kivu long-haired shrew (C. lanosa)
  • Ussuri white-toothed shrew (C. lasiura)
  • Latona's shrew (C. latona)
  • Sulawesi shrew (C. lea)
  • Sumatran giant shrew (C. lepidura)
  • Bicolored shrew (C. leucodon)
  • Sulawesi tiny shrew (C. levicula)
  • Butiaba naked-tailed shrew (C. littoralis)
  • Savanna swamp shrew (C. longipes)
  • Lucina's shrew (C. lucina)
  • Ludia's shrew (C. ludia)
  • Moonshine shrew (C. luna)
  • Mauritanian shrew (C. lusitania)
  • MacArthur's shrew (C. macarthuri)
  • MacMillan's shrew (C. macmillani)
  • Nyiro shrew (C. macowi)
  • Malayan shrew (C. malayana)
  • Manenguba shrew (C. manengubae)
  • Makwassie musk shrew (C. maquassiensis)
  • Swamp musk shrew (C. mariquensis)
  • Gracile naked-tailed shrew (C. maurisca)
  • Javanese shrew (C. maxi)
  • Mindoro shrew (C. mindorus)
  • Sri Lankan long-tailed shrew (C. miya)
  • Kilimanjaro shrew (C. monax)
  • Sunda shrew (C. monticola)
  • Montane white-toothed shrew (C. montis)
  • West African long-tailed shrew (C. muricauda)
  • Mossy forest shrew (C. musseri)
  • Ugandan musk shrew (C. mutesae)
  • Somali dwarf shrew (C. nana)
  • Savanna dwarf shrew (C. nanilla)
  • Peninsular shrew (C. negligens)
  • Negros shrew (C. negrina)
  • Nicobar shrew (C. nicobarica)
  • Nigerian shrew (C. nigeriae)
  • Blackish white-toothed shrew (C. nigricans)
  • Black-footed shrew (C. nigripes)
  • African black shrew (C. nigrofusca)
  • Nimba shrew (C. nimbae)
  • Niobe's shrew (C. niobe)
  • West African pygmy shrew (C. obscurior)
  • African giant shrew (C. olivieri)
  • Oriental shrew (C. orientalis)
  • Ryukyu shrew (C. orii)
  • Palawan shrew (C. palawanensis)
  • Panay shrew (C. panayensis)
  • Sumatran long-tailed shrew (C. paradoxura)
  • Small-footed shrew (C. parvipes)
  • Sahelian tiny shrew (C. pasha)
  • Pale gray shrew (C. pergrisea)
  • Guramba shrew (C. phaeura)
  • C. phanluongi
  • Phu Hoc shrew (C. phuquocensis)
  • Cameroonian shrew (C. picea)
  • Pitman's shrew (C. pitmani)
  • Flat-headed shrew (C. planiceps)
  • Fraser's musk shrew (C. poensis)
  • Polia's shrew (C. polia)
  • Kashmir white-toothed shrew (C. pullata)
  • Rainey's shrew (C. raineyi)
  • Negev shrew (C. ramona)
  • Chinese white-toothed shrew (C. rapax)
  • Egyptian pygmy shrew (C. religiosa)
  • Sulawesi white-handed shrew (C. rhoditis)
  • Roosevelt's shrew (C. roosevelti)
  • Greater white-toothed shrew (C. russula)
  • Ugandan lowland shrew (C. selina)
  • Lesser rock shrew (C. serezkyensis)
  • Asian lesser white-toothed shrew (C. shantungensis)
  • Siberian shrew (C. sibirica)
  • Sicilian shrew (C. sicula)
  • Lesser gray-brown musk shrew (C. silacea)
  • Desert musk shrew (C. smithii)
  • Sokolov's shrew (C. sokolovi)
  • Somali shrew (C. somalica)
  • Kahuzi swamp shrew (C. stenocephala)
  • Lesser white-toothed shrew (C. suaveolens)
  • Iranian shrew (C. susiana)
  • Tanzanian shrew (C. tansaniana)
  • Tarella shrew (C. tarella)
  • Saharan shrew (C. tarfayensis)
  • Telford's shrew (C. telfordi)
  • Timor shrew (C. tenuis)
  • Thalia's shrew (C. thalia)
  • Therese's shrew (C. theresae)
  • São Tomé shrew (C. thomensis)
  • Christmas Island shrew (C. trichura)
  • Turbo shrew (C. turba)
  • Ultimate shrew (C. ultima)
  • Usambara shrew (C. usambarae)
  • Savanna path shrew (C. viaria)
  • Mamfe shrew (C. virgata)
  • Voi shrew (C. voi)
  • Voracious shrew (C. vorax)
  • Banka shrew (C. vosmaeri)
  • Lesser Ryukyu shrew (C. watasei)
  • Whitaker's shrew (C. whitakeri)
  • Wimmer's shrew (C. wimmeri)
  • Hainan Island shrew (C. wuchihensis)
  • Xanthippe's shrew (C. xantippe)
  • Yankari shrew (C. yankariensis)
  • Mikhail Zaitsev's shrew (C. zaitsevi )
  • Zaphir's shrew (C. zaphiri)
  • Zarudny's rock shrew (C. zarudnyi)
  • Upemba shrew (C. zimmeri)
  • Cretan shrew (C. zimmermanni)
Diplomesodon
  • Piebald shrew (D. pulchellus)
Feroculus
  • Kelaart's long-clawed shrew (F. feroculus)
Palawanosorex
  • Palawan moss shrew (P. muscorum)
Paracrocidura
(Large-headed
shrews)
  • Grauer's large-headed shrew (P. graueri)
  • Greater large-headed shrew (P. maxima)
  • Lesser large-headed shrew (P. schoutedeni)
Ruwenzorisorex
  • Ruwenzori shrew (R. suncoides)
Scutisorex
  • Hero shrew (S. somereni)
  • Thor's hero shrew (S. thori)
Solisorex
  • Pearson's long-clawed shrew (S. pearsoni)
Suncus
  • Taita shrew (S. aequatorius)
  • Black shrew (S. ater)
  • Day's shrew (S. dayi)
  • Etruscan shrew (S. etruscus)
  • Sri Lankan shrew (S. fellowesgordoni)
  • Bornean pygmy shrew (S. hosei)
  • Least dwarf shrew (S. infinitesimus)
  • Greater dwarf shrew (S. lixus)
  • Madagascan pygmy shrew (S. madagascariensis)
  • Malayan pygmy shrew (S. malayanus)
  • Climbing shrew (S. megalura)
  • Flores shrew (S. mertensi)
  • Asian highland shrew (S. montanus)
  • Asian house shrew (S. murinus)
  • Remy's pygmy shrew (S. remyi)
  • Anderson's shrew (S. stoliczkanus)
  • Lesser dwarf shrew (S. varilla)
  • Jungle shrew (S. zeylanicus)
Sylvisorex
(Forest shrews)
  • Dudu Akaibe's pygmy shrew (S. akaibei)
  • Cameroonian forest shrew (S. cameruniensis)
  • Corbet's forest shrew (S. corbeti)
  • Grant's forest shrew (S. granti)
  • Howell's forest shrew (S. howelli)
  • Bioko forest shrew (S. isabellae)
  • Johnston's forest shrew (S. johnstoni)
  • Kongana shrew (S. konganensis)
  • Moon forest shrew (S. lunaris)
  • Mount Cameroon forest shrew (S. morio)
  • Greater forest shrew (S. ollula)
  • Lesser forest shrew (S. oriundus)
  • Rain forest shrew (S. pluvialis)
  • Volcano shrew (S. vulcanorum)
Anourosoricini
Anourosorex
(Asian mole shrews)
  • Assam mole shrew (A. assamensis)
  • Giant mole shrew (A. schmidi)
  • Chinese mole shrew (A. squamipes)
  • Taiwanese mole shrew (A. yamashinai)
Blarinellini
Blarinella
(Asiatic short-tailed
shrews)
  • Indochinese short-tailed shrew (B. griselda)
  • Asiatic short-tailed shrew (B. quadraticauda)
  • Burmese short-tailed shrew (B. wardi)
Blarinini
Blarina
(American short-tailed
shrews)
  • Northern short-tailed shrew (B. brevicauda)
  • Southern short-tailed shrew (B. carolinensis)
  • Elliot's short-tailed shrew (B. hylophaga)
  • Everglades short-tailed shrew (B. peninsulae)
  • Sherman's short-tailed shrew (B. shermani)
Cryptotis
(Small-eared
shrews)
C. mexicana group
Mexican small-eared shrew (C. mexicana)
Nelson's small-eared shrew (C. nelsoni)
Grizzled Mexican small-eared shrew (C. obscura)
Phillips' small-eared shrew (C. phillipsii)
C. goldmani set
Central Mexican broad-clawed shrew (C. alticola)
Goldman's broad-clawed shrew (C. goldmani)
Goodwin's broad-clawed shrew (C. goodwini)
Guatemalan broad-clawed shrew (C. griseoventris)
C. lacertosus
C. mam
Oaxacan broad-clawed shrew (C. peregrina)
C. nigrescens group
Eastern Cordillera small-footed shrew (C. brachyonyx)
Colombian small-eared shrew (C. colombiana)
Honduran small-eared shrew (C. hondurensis)
Yucatan small-eared shrew (C. mayensis)
Darién small-eared shrew (C. mera)
Merriam's small-eared shrew (C. merriami)
Blackish small-eared shrew (C. nigrescens)
C. thomasi group
Southern Colombian small-eared shrew (C. andinus)
Ecuadorian small-eared shrew (C. equatoris)
Rainer's small-eared shrew (C. huttereri)
Medellín small-eared shrew (C. medellinia)
Merida small-eared shrew (C. meridensis)
Wandering small-eared shrew (C. montivaga)
Peruvian small-eared shrew (C. peruviensis)
Scaly-footed small-eared shrew (C. squaipes)
Tamá small-eared shrew (C. tamensis)
Thomas's small-eared shrew (C. thomasi)
C. parva group
Central American least shrew (C. orophila)
North American least shrew (C. parva)
Tropical small-eared shrew (C. tropicalis)
Ungrouped / relict
Enders's small-eared shrew (C. endersi)
Talamancan small-eared shrew (C. gracilis)
Big Mexican small-eared shrew (C. magna)
Nectogalini
Chimarrogale
(Asiatic water
shrews)
  • Malayan water shrew (C. hantu)
  • Himalayan water shrew (C. himalayica)
  • Bornean water shrew (C. phaeura)
  • Japanese water shrew (C. platycephalus)
  • Chinese water shrew (C. styani)
  • Sumatran water shrew (C. sumatrana)
Chodsigoa
  • Van Sung's shrew (C. caovansunga)
  • De Winton's shrew (C. hypsibia)
  • Lamulate shrew (C. lamula)
  • Lowe's shrew (C. parca)
  • Pygmy brown-toothed shrew (C. parva)
  • Salenski's shrew (C. salenskii)
  • Smith's shrew (C. smithii)
  • Lesser Taiwanese shrew (C. sodalis)
Episoriculus
  • Hodgsons's brown-toothed shrew (E. caudatus)
  • Taiwanese brown-toothed shrew (E. fumidus)
  • Long-tailed brown-toothed shrew (E. leucops)
  • Long-tailed mountain shrew (E. macrurus)
Nectogale
  • Elegant water shrew (N. elegans)
Neomys
  • Mediterranean water shrew (N. anomalus)
  • Eurasian water shrew (N. fodiens)
  • Transcaucasian water shrew (N. teres)
Soriculus
  • Himalayan shrew (S. nigrescens)
Notiosoricini
Megasorex
  • Mexican shrew (M. gigas)
Notiosorex
  • Cockrum's gray shrew (N. cockrumi)
  • Crawford's gray shrew (N. crawfordi)
  • Large-eared gray shrew (N. evotis)
  • Villa's gray shrew (N. villai)
Sorex
(Long-tailed
shrews)
Subgenus
Otisorex
  • Long-tailed shrew (S. dispar)
  • Smoky shrew (S. fumeus)
  • American pygmy shrew (S. hoyi)
  • Large-toothed shrew (S. macrodon)
  • Carmen mountain shrew (S. milleri)
  • Dwarf shrew (S. nanus)
  • Mexican long-tailed shrew (S. oreopolus)
  • Orizaba long-tailed shrew (S. orizabae)
  • Ornate shrew (S. ornatus)
  • Inyo shrew (S. tenellus)
  • Verapaz shrew (S. veraepacis)
S. vagrans complex
Glacier Bay water shrew (S. alaskanus)
Baird's shrew (S. bairdii)
Marsh shrew (S. bendirii)
Montane shrew (S. monticolus)
New Mexico shrew (S. neomexicanus)
Pacific shrew (S. pacificus)
American water shrew (S. palustris)
Fog shrew (S. sonomae)
Vagrant shrew (S. vagrans)
S. cinereus group
Kamchatka shrew (S. camtschatica)
Cinereus shrew (S. cinereus)
Prairie shrew (S. haydeni)
Saint Lawrence Island shrew (S. jacksoni)
Paramushir shrew (S. leucogaster)
Southeastern shrew (S. longirostris)
Mount Lyell shrew (S. lyelli)
Portenko's shrew (S. portenkoi)
Preble's shrew (S. preblei)
Pribilof Island shrew (S. pribilofensis)
Olympic shrew (S. rohweri)
Barren ground shrew (S. ugyunak)
Subgenus
Sorex
  • Dneper common shrew (S. averini)
  • Lesser striped shrew (S. bedfordiae)
  • Greater stripe-backed shrew (S. cylindricauda)
  • Chinese highland shrew (S. excelsus)
  • Azumi shrew (S. hosonoi)
  • Chinese shrew (S. sinalis)
  • Alaska tiny shrew (S. yukonicus)
S. alpinus group
Alpine shrew (S. alpinus)
Ussuri shrew (S. mirabilis)
S. araneus group
Valais shrew (S. antinorii)
Common shrew (S. araneus)
Udine shrew (S. arunchi)
Crowned shrew (S. coronatus)
Siberian large-toothed shrew (S. daphaenodon)
Iberian shrew (S. granarius)
Caucasian shrew (S. satunini)
S. arcticus group
Arctic shrew (S. arcticus)
Maritime shrew (S. maritimensis)
S. tundrensis group
Tien Shan shrew (S. asper)
Gansu shrew (S. cansulus)
Tundra shrew (S. tundrensis)
S. minutus group
Buchara shrew (S. buchariensis)
Kozlov's shrew (S. kozlovi)
Caucasian pygmy shrew (S. volnuchini)
S. caecutiens group
Laxmann's shrew (S. caecutiens)
Taiga shrew (S. isodon)
Eurasian least shrew (S. minutissimus)
Eurasian pygmy shrew (S. minutus)
Flat-skulled shrew (S. roboratus)
Shinto shrew (S. shinto)
Long-clawed shrew (S. unguiculatus)
S. gracillimus group
Slender shrew (S. gracillimus)
S. raddei group
Radde's shrew (S. raddei)
S. samniticus group
Apennine shrew (S. samniticus)
incertae sedis
  • Arizona shrew (S. arizonae)
  • Zacatecas shrew (S. emarginatus)
  • Merriam's shrew (S. merriami)
  • Kashmir pygmy shrew (S. planiceps)
  • Saussure's shrew (S. saussurei)
  • Sclater's shrew (S. sclateri)
  • San Cristobal shrew (S. stizodon)
  • Tibetan shrew (S. thibetanus)
  • Trowbridge's shrew (S. trowbridgii)
  • Chestnut-bellied shrew (S. ventralis)
  • Veracruz shrew (S. veraecrucis)
Congosorex
(Congo shrews)
  • Phillips's shrew (C. phillipsorum)
  • Greater Congo shrew (C. polli)
  • Lesser Congo shrew (C. verheyeni)
Myosorex
(Mouse shrews)
  • Babault's mouse shrew (M. babaulti)
  • Montane mouse shrew (M. blarina)
  • Bururi forest shrew (M. bururiensis)
  • Dark-footed mouse shrew (M. cafer)
  • Eisentraut's mouse shrew (M. eisentrauti)
  • Geata mouse shrew (M. geata)
  • Nyika mouse shrew (M. gnoskei)
  • Kihaule's mouse shrew (M. kihaulei)
  • Long-tailed forest shrew (M. longicaudatus)
  • Oku mouse shrew (M. okuensis)
  • Rumpi mouse shrew (M. rumpii)
  • Schaller's mouse shrew (M. schalleri)
  • Sclater's mouse shrew (M. sclateri)
  • Thin mouse shrew (M. tenuis)
  • Forest shrew (M. varius)
  • Kilimanjaro mouse shrew (M. zinki)
Surdisorex
(African mole
shrews)
  • Aberdare mole shrew (S. norae)
  • Mount Kenya mole shrew (S. polulus)
Scalopinae
(New World moles
and relatives)
Condylura
  • Star-nosed mole (C. cristata)
Parascalops
  • Hairy-tailed mole (P. breweri)
Scalopus
  • Eastern mole (S. aquaticus)
Scapanulus
  • Gansu mole (S. oweni)
Scapanus
(Western North
American moles)
  • Broad-footed mole (S. latimanus)
  • Coast mole (S. orarius)
  • Townsend's mole (S. townsendii)
Talpinae
(Old World moles
and relatives)
Desmana
  • Russian desman (D. moschata)
Dymecodon
  • True's shrew mole (D. pilirostris)
Euroscaptor
  • Greater Chinese mole (E. grandis)
  • Kloss's mole (E. klossi)
  • Long-nosed mole (E. longirostris)
  • Himalayan mole (E. micrura)
  • Small-toothed mole (E. parvidens)
  • Vietnamese mole (E. subanura)
Galemys
  • Pyrenean desman (G. pyrenaicus)
Mogera
  • Echigo mole (M. etigo)
  • Insular mole (M. insularis)
  • Kano mole (M. kanoana)
  • Kobe mole (M. kobeae)
  • Small Japanese mole (M. imaizumii)
  • Large mole (M. robusta)
  • Sado mole (M. tokudae)
  • Japanese mole (M. wogura)
  • Senkaku mole (M. uchidai)
Neurotrichus
  • American shrew mole (N. gibbsii)
Oreoscaptor
  • Japanese mountain mole (O. mizura)
Parascaptor
  • White-tailed mole (P. leucura)
Scaptochirus
  • Short-faced mole (S. moschatus)
Scaptonyx
  • Long-tailed mole (S. fusicaudus)
Talpa
  • Altai mole (T. altaica)
  • Aquitanian mole (T. aquitania)
  • Blind mole (T. caeca)
  • Caucasian mole (T. caucasica)
  • European mole (T. europaea)
  • Père David's mole (T. davidiana)
  • Levant mole (T. levantis)
  • Martino's mole (T. martinorum)
  • Spanish mole (T. occidentalis)
  • Ognev's mole (T. ognevi)
  • Roman mole (T. romana)
  • Balkan mole (T. stankovici)
  • Talysch mole (T. talyschensis)
Urotrichus
  • Japanese shrew mole (U. talpoides)
Uropsilinae
(Chinese shrew-like
moles)
  • Equivalent-teeth shrew mole (U. aequodonenia)
  • Anderson's shrew mole (U. andersoni)
  • Black-backed shrew mole (U. atronates)
  • Dabie Mountains shrew mole (U. dabieshanensis)
  • Gracile shrew mole (U. gracilis)
  • Inquisitive shrew mole (U. investigator)
  • Snow Mountain shrew mole (U. nivatus)
  • Chinese shrew mole (U. soricipes)
Atopogale
  • Cuban solenodon (A. cubana)
Solenodon
  • Hispaniolan solenodon (S. paradoxus)
Taxon identifiers
Galericinae