Macrobaenidae

Extinct family of turtles

Macrobaenidae Temporal range: Early Cretaceous–Paleocene PreꞒ Ꞓ O S D C P T J K Pg N

Osteopygis emarginatus carapace
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Pantestudines
Clade:	Testudinata
Clade:	Perichelydia
Family:	†Macrobaenidae Sukhanov, 1964
Genera
See text

Macrobaenidae is an extinct family of turtles, known from the Early Cretaceous to Paleogene of Laurasia. Their relationships to other turtles and whether they form a monophlyletic group are controversial. They are typically interpreted as stem or crown group cryptodires, but some more recent analyses have found them to lie outside crown group Testudines. Macrobaenids can be distinguished from other testudinatans by the presence of a carotid fenestra, cruciform plastron with strap-like epiplastra, and a lack of extragulars.^[1]

Genera

Anatolemys^[2] Central Asia, Late Cretaceous ( Khodzhakul Formation, Uzbekistan, Cenomanian, Bissekty Formation, Uzbekistan, Turonian, Bostobe Formation, Kazakhstan, Santonian Yalovach Formation, Tajikistan, Santonian)
Appalachemys^[3] Mooreville Chalk, Alabama, Late Cretaceous (Santonian–Campanian)
Asiachelys^[2] Khulsangol Formation, Mongolia, Early Cretaceous (Albian)
Aurorachelys^[2] Kanguk Formation, Canada,Turonian
Changmachelys Xiagou Formation, China, Early Cretaceous (Aptian)
Kirgizemys^[2] Murtoi Formation, Khilok Formation, Ilek Formation, Russia, Early Cretaceous (Barremian-Aptian), Alamyshik Formation, Kyrgyzstan, Albian, Geoncheonri Formation, South Korea, Albian Khodzhakul Formation, Uzbekistan, Cenomanian
Judithemys North America, Late Cretaceous–Paleocene
Gallica^[4] Paris Basin, France, Paleocene
Macrobaena^[2] Naran Bulak Formation, Mongolia, Paleocene
Osteopygis^[2] Late Cretaceous-Paleocene, North America
Oxemys^[2] Khodzhakul Formation, Uzbekistan, Cenomanian
Yakemys Phu Kradung Formation, Thailand, Early Cretaceous (Berriasian)

Hongkongochelys from the Middle-Late Jurassic of China has sometimes been attributed to the family,^[4] but other times has been attriuted to Sinemydidae, a group which has an unresolved relationship with Macrobaenidae.

References

^ Joyce, Walter G.; Anquetin, Jérémy; Cadena, Edwin-Alberto; Claude, Julien; Danilov, Igor G.; Evers, Serjoscha W.; Ferreira, Gabriel S.; Gentry, Andrew D.; Georgalis, Georgios L.; Lyson, Tyler R.; Pérez-García, Adán (December 2021). "A nomenclature for fossil and living turtles using phylogenetically defined clade names". Swiss Journal of Palaeontology. 140 (1): 5. doi:10.1186/s13358-020-00211-x. hdl:11336/155192. ISSN 1664-2376.
^ ^a ^b ^c ^d ^e ^f ^g "Macrobaenidae". FossilWorks. Retrieved 17 December 2021.
^ Gentry, A. D.; Kiernan, C. R.; Parham, J. F. (2022). "A large non-marine turtle from the Upper Cretaceous of Alabama and a review of North American "Macrobaenids"". The Anatomical Record. doi:10.1002/ar.25054.
^ ^a ^b Adán Pérez-García (2020). "A European Cenozoic 'Macrobaenid:' New Data about the Paleocene Arrival of Several Turtle Lineages to Europe". Journal of Vertebrate Paleontology. 40 (4): e1795874. doi:10.1080/02724634.2020.1795874. S2CID 225151817.

Taxon identifiers
Macrobaenidae	Wikidata: Q12071605 GBIF: 4819118 IRMNG: 117524 Open Tree of Life: 3606561 Paleobiology Database: 56472

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Suborder

Superfamily

Family

Genus

Cryptodira

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.