Malagasy civet

Species of carnivore

Malagasy civet
at Ranomafana National Park
Conservation status

Vulnerable  (IUCN 3.1)[1]
CITES Appendix II (CITES)[2]
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Eupleridae
Subfamily: Euplerinae
Genus: Fossa
Gray, 1864[3]
Species:
F. fossana
Binomial name
Fossa fossana
(Müller, 1776)[3]
Malagasy civet range[4]
Synonyms

Viverra fossana Müller, 1776

The Malagasy or striped civet (Fossa fossana), also known as the fanaloka (Malagasy, [fə̥ˈnaluk]) or jabady,[5] is an euplerid endemic to Madagascar.[6] It is the only species in genus Fossa.

The Malagasy civet is a small mammal, about 47 centimetres (19 in) long excluding the tail (which is only about 20 centimetres (7.9 in)). It can weigh 1.5 to 2.0 kilograms (3.3 to 4.4 lb). It is endemic to the tropical forests of Madagascar. Malagasy civets are nocturnal. It eats small vertebrates, insects, aquatic animals, and eggs stolen from birds' nests. The mating season of the Malagasy civet is August to September and the gestation period is three months, ending with the birth of one young. The Malagasy civet is listed as Vulnerable by International Union for Conservation of Nature (IUCN).[1]

The Malagasy name fanaloka is related to the Malay word pelanduk "mousedeer" (via metathesis) likely due to their similar sizes and silhouette.[7]

Classification

The Malagasy civet was to be placed in the subfamily Hemigalinae with the banded palm civets and then in its own subfamily, Fossinae, because of similarities with others in the group pointed out by Gregory, but it is now classified as a member of the subfamily Euplerinae, after Pocock pointed out more similarities with that one.[8]

Description

The Malagasy civet is a small mammal, about 47 centimetres (19 in) long excluding the tail (which is only about 20 centimetres (7.9 in)). The males can weigh up to 1.9 kilograms (4.2 lb), and the females can weigh up to 1.75 kilograms (3.9 lb). It is the second largest carnivore in Madagascar after the fossa.[9][10] it may be confused with the small Indian civet (Viverricula indica).[11] It has a short coat greyish beige or brown in colour, with dark black horizontal stripes running from head to tail, where the stripes are vertical, wrapping around the bushier tail. The stripes morph into spots near the belly. Its legs are short and very thin.[11]

Behavior

The Malagasy civet is nocturnal, though sources disagree over whether it is solitary or, unusual among euplerids, lives in pairs. It is not a good climber and frequents ravines. It eats small vertebrates (mammals, small birds, reptiles, and amphibians), insects, aquatic animals, and eggs stolen from birds' nests.[12][13] It is shy and secretive.[14] Their vocalizations are similar to crying and groaning, as well as a sound similar to coq-coq. Pairs of males and females defend a large area (around 50 hectares (120 acres)) as their territory.[15] In the winter, it may store fat in its tail, which can make up 25% of their weight.[9][16] The mating season of the Malagasy civet is August to September and the gestation period is around three months, ending with the birth of one young. The young are rather well-developed, weigh around 65 to 70 grams (2.3 to 2.5 oz), and are weaned in two to three months, leaving their parents at around one year old. The average lifespan of a Malagasy civet is about 21 years in captivity.[1][17][18]

Distribution and habitat

The Malagasy civet is found in lowland and rainforest areas of Eastern and Northern areas of Madagascar, and can also be found in humid and isolated forests in Amber Mountain National Park, and farther north in the less-humid forests of Ankarana Reserve. It can be found from sea level to 1,600 metres (5,200 ft) above sea level, but is only common up to 1,000 metres (3,300 ft) above sea level.[1][11]

Conservation status

The Malagasy civet is listed as Vulnerable by International Union for Conservation of Nature (IUCN), with a decreasing population. Though threatened by deforestation, hunting, charcoal production, logging, and competition from introduced species such as dogs, cats, and small Indian civets, it is locally common. Introduced animals such as dogs are likely to prey on Malagasy civets.[1] Its range is now reduced to isolated patches.[18]

References

  1. ^ a b c d e Hawkins, F. (2015). "Fossa fossana". IUCN Red List of Threatened Species. 2015: e.T8668A45197868. doi:10.2305/IUCN.UK.2015-4.RLTS.T8668A45197868.en. Retrieved 12 November 2021.
  2. ^ "Appendices | CITES". cites.org. Retrieved 2022-01-14.
  3. ^ a b "Fossa fossana". ITIS. Retrieved 1 January 2015.
  4. ^ Hawkins, A. F .A. (2008). "Fossa fossana". IUCN Red List of Threatened Species. 2008. Retrieved 22 March 2009.old-form url
  5. ^ Nick Garbutt; Hilary Bradt; Derek Schuurman (2008). Madagascar Wildlife. Bradt Travel Guides. p. 78. ISBN 978-1-84162-245-3.
  6. ^ Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 560. ISBN 978-0-8018-8221-0. OCLC 62265494.
  7. ^ Blench, Roger M.; Walsh, Martin (2009). Faunal names in Malagasy: their etymologies and implications for the prehistory of the East African coast (PDF). Eleventh International Conference on Austronesian Linguistics (11 ICAL). Aussois, France. pp. 1–31.
  8. ^ Anjali Goswami; Anthony Friscia (29 July 2010). Carnivoran Evolution: New Views on Phylogeny, Form and Function. Cambridge University Press. pp. 68–70. ISBN 978-1-139-48853-2.
  9. ^ a b "Fossa fossana". animaldiversity.org. Retrieved 1 January 2015.
  10. ^ Steven Roger Fischer (15 February 2013). Islands: From Atlantis to Zanzibar. Reaktion Books. p. 66. ISBN 978-1-78023-053-5.
  11. ^ a b c Nick Garbutt (2007). Mammals of Madagascar: A Complete Guide. Yale University Press. p. 209. ISBN 978-0-300-12550-4.
  12. ^ Richard T. Corlett; Richard B. Primack (3 March 2011). Tropical Rain Forests: An Ecological and Biogeographical Comparison. John Wiley & Sons. p. 136. ISBN 978-1-4443-9228-9.
  13. ^ "Fossa fossana (Malagasy civet)". Animal Diversity Web.
  14. ^ Hilary Bradt; Daniel Austin (1 July 2014). Madagascar. Bradt Travel Guides. p. 66. ISBN 978-1-84162-498-3.
  15. ^ Daniel Austin (10 November 2014). Madagascar Wildlife. Bradt Travel Guides. p. 84. ISBN 978-1-84162-557-7.
  16. ^ Animals: A Visual Encyclopedia (Second Edition): A Visual Encyclopedia. DK Publishing. 19 March 2012. p. 83. ISBN 978-0-7566-9896-6.
  17. ^ "Fossa fossana". Encyclopedia of Life. Retrieved 1 January 2015.
  18. ^ a b Ronald M. Nowak (2005). Walker's Carnivores of the World. JHU Press. pp. 201–202. ISBN 978-0-8018-8032-2.

Sources

  • Macdonald, David (ed). The Encyclopedia of Mammals. (New York, 1984)
  • Anderson, Simon (ed). Simon & Schuster's Guide to Mammals. (Milan, 1982)

External links

Wikispecies has information related to Fossa fossana.
Wikimedia Commons has media related to Fossa_fossana.
  • ARKive - images and movies of the striped civet (Fossa fossana)
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Extant Carnivora species
Suborder Feliformia
Nandiniidae
Nandinia
  • African palm civet (N. binotata)
Herpestidae
(Mongooses)
Atilax
  • Marsh mongoose (A. paludinosus)
Bdeogale
  • Bushy-tailed mongoose (B. crassicauda)
  • Jackson's mongoose (B. jacksoni)
  • Black-footed mongoose (B. nigripes)
Crossarchus
  • Alexander's kusimanse (C. alexandri)
  • Angolan kusimanse (C. ansorgei)
  • Common kusimanse (C. obscurus)
  • Flat-headed kusimanse (C. platycephalus)
Cynictis
  • Yellow mongoose (C. penicillata)
Dologale
  • Pousargues's mongoose (D. dybowskii)
Helogale
  • Ethiopian dwarf mongoose (H. hirtula)
  • Common dwarf mongoose (H. parvula)
Herpestes
  • Angolan slender mongoose (H. flavescens)
  • Egyptian mongoose (H. ichneumon)
  • Somalian slender mongoose (H. ochracea)
  • Cape gray mongoose (H. pulverulenta)
  • Common slender mongoose (H. sanguinea)
Ichneumia
  • White-tailed mongoose (I. albicauda)
Liberiictus
  • Liberian mongoose (L. kuhni)
Mungos
  • Gambian mongoose (M. gambianus)
  • Banded mongoose (M. mungo)
Paracynictis
  • Selous's mongoose (P. selousi)
Rhynchogale
  • Meller's mongoose (R. melleri)
Suricata
  • Meerkat (S. suricatta)
Urva
  • Small Indian mongoose (U. auropunctata)
  • Short-tailed mongoose (U. brachyura)
  • Indian grey mongoose (U. edwardsii)
  • Indian brown mongoose (U. fusca)
  • Javan mongoose (U. javanica)
  • Collared mongoose (U. semitorquata)
  • Ruddy mongoose (U. smithii)
  • Crab-eating mongoose (U. urva)
  • Stripe-necked mongoose (U. vitticolla)
Xenogale
  • Long-nosed mongoose (X. naso)
Hyaenidae
(Hyenas)
Crocuta
  • Spotted hyena (C. crocuta)
Hyaena
  • Striped hyena (H. hyaena)
Parahyaena
  • Brown hyena (P. brunnea)
Proteles
  • Aardwolf (P. cristata)
Felidae
Large family listed below
Viverridae
Large family listed below
Eupleridae
Small family listed below
Family Felidae
Felinae
Acinonyx
  • Cheetah (A. jubatus)
Caracal
  • African golden cat (C. aurata)
  • Caracal (C. caracal)
Catopuma
  • Bay cat (C. badia)
  • Asian golden cat (C. temminckii)
Felis
  • Chinese mountain cat (F. bieti)
  • Domestic cat (F. catus)
  • Jungle cat (F. chaus)
  • African wildcat (F. lybica)
  • Sand cat (F. margarita)
  • Black-footed cat (F. nigripes)
  • European wildcat (F. silvestris)
Herpailurus
  • Jaguarundi (H. yagouaroundi)
Leopardus
  • Pampas cat (L. colocola)
  • Geoffroy's cat (L. geoffroyi)
  • Kodkod (L. guigna)
  • Southern tiger cat (L. guttulus)
  • Andean mountain cat (L. jacobita)
  • Ocelot (L. pardalis)
  • Oncilla (L. tigrinus)
  • Margay (L. wiedii)
Leptailurus
  • Serval (L. serval)
Lynx
  • Canada lynx (L. canadensis)
  • Eurasian lynx (L. lynx)
  • Iberian lynx (L. pardinus)
  • Bobcat (L. rufus)
Otocolobus
  • Pallas's cat (O. manul)
Pardofelis
  • Marbled cat (P. marmorata)
Prionailurus
  • Leopard cat (P. bengalensis)
  • Sunda leopard cat (P. javanensis)
  • Flat-headed cat (P. planiceps)
  • Rusty-spotted cat (P. rubiginosus)
  • Fishing cat (P. viverrinus)
Puma
  • Cougar (P. concolor)
Pantherinae
Panthera
  • Lion (P. leo)
  • Jaguar (P. onca)
  • Leopard (P. pardus)
  • Tiger (P. tigris)
  • Snow leopard (P. uncia)
Neofelis
  • Sunda clouded leopard (N. diardi)
  • Clouded leopard (N. nebulosa)
Prionodon
  • Banded linsang (P. linsang)
  • Spotted linsang (P. pardicolor)
Family Viverridae
Paradoxurinae
Arctictis
  • Binturong (A. binturong)
Arctogalidia
  • Small-toothed palm civet (A. trivirgata)
Macrogalidia
  • Sulawesi palm civet (M. musschenbroekii)
Paguma
  • Masked palm civet (P. larvata)
Paradoxurus
  • Asian palm civet (P. hermaphroditus)
  • Brown palm civet (P. jerdoni)
  • Golden palm civet (P. zeylonensis)
Hemigalinae
Chrotogale
  • Owston's palm civet (C. owstoni)
Cynogale
  • Otter civet (C. bennettii)
Diplogale
  • Hose's palm civet (D. hosei)
Hemigalus
  • Banded palm civet (H. derbyanus)
Viverrinae
Civettictis
  • African civet (C. civetta)
Viverra
  • Malabar large-spotted civet (V. civettina)
  • Large-spotted civet (V. megaspila)
  • Malayan civet (V. tangalunga)
  • Large Indian civet (V. zibetha)
Viverricula
  • Small Indian civet (V. indica)
Genettinae
Genetta
(Genets)
  • Abyssinian genet (G. abyssinica)
  • Angolan genet (G. angolensis)
  • Bourlon's genet (G. bourloni)
  • Crested servaline genet (G. cristata)
  • Common genet (G. genetta)
  • Johnston's genet (G. johnstoni)
  • Letaba genet (G. letabae)
  • Rusty-spotted genet (G. maculata)
  • Pardine genet (G. pardina)
  • Aquatic genet (G. piscivora)
  • King genet (G. poensis)
  • Servaline genet (G. servalina)
  • Hausa genet (G. thierryi)
  • Cape genet (G. tigrina)
  • Giant forest genet (G. victoriae)
  • South African small-spotted genet (G. felina)
Poiana
  • Central African oyan (P. richardsonii)
  • West African oyan (P. leightoni)
Family Eupleridae
Euplerinae
Cryptoprocta
  • Fossa (C. ferox)
Eupleres
  • Eastern falanouc (E. goudotii)
  • Western falanouc (E. major)
Fossa
  • Malagasy civet (F. fossana)
Galidiinae
Galidia
  • Ring-tailed vontsira (G. elegans)
Galidictis
  • Broad-striped Malagasy mongoose (G. fasciata)
  • Grandidier's mongoose (G. grandidieri)
Mungotictis
  • Narrow-striped mongoose (M. decemlineata)
Salanoia
  • Brown-tailed mongoose (S. concolor)
  • Durrell's vontsira (S. durrelli)
Suborder Caniformia (cont. below)
Ursidae
(Bears)
Ailuropoda
  • Giant panda (A. melanoleuca)
Helarctos
  • Sun bear (H. malayanus)
Melursus
  • Sloth bear (M. ursinus)
Tremarctos
  • Spectacled bear (T. ornatus)
Ursus
  • American black bear (U. americanus)
  • Brown bear (U. arctos)
  • Polar bear (U. maritimus)
  • Asian black bear (U. thibetanus)
Mephitidae
(Skunks)
Conepatus
(Hog-nosed
skunks)
  • Molina's hog-nosed skunk (C. chinga)
  • Humboldt's hog-nosed skunk (C. humboldtii)
  • American hog-nosed skunk (C. leuconotus)
  • Striped hog-nosed skunk (C. semistriatus)
Mephitis
  • Hooded skunk (M. macroura)
  • Striped skunk (M. mephitis)
Mydaus
  • Sunda stink badger (M. javanensis)
  • Palawan stink badger (M. marchei)
Spilogale
(Spotted skunks)
  • Southern spotted skunk (S. angustifrons)
  • Western spotted skunk (S. gracilis)
  • Eastern spotted skunk (S. putorius)
  • Pygmy spotted skunk (S. pygmaea)
Procyonidae
(Raccoons, coatis, olingos)
Bassaricyon
(Olingos)
  • Eastern lowland olingo (B. alleni)
  • Northern olingo (B. gabbii)
  • Western lowland olingo (B. medius)
  • Olinguito (B. neblina)
Bassariscus
  • Ringtail (B. astutus)
  • Cacomistle (B. sumichrasti)
Nasua
(Coatis inclusive)
  • White-nosed coati (N. narica)
  • South American coati (N. nasua)
Nasuella
(Coatis inclusive)
  • Eastern mountain coati (N. meridensis)
  • Western mountain coati (N. olivacea)
Potos
  • Kinkajou (P. flavus)
Procyon
  • Crab-eating raccoon (P. cancrivorus)
  • Raccoon (P. lotor)
  • Cozumel raccoon (P. pygmaeus)
Ailuridae
Ailurus
  • Red panda (A. fulgens)
Suborder Caniformia (cont. above)
Otariidae
(Eared seals)
(includes fur seals
and sea lions)

(Pinniped inclusive)
Arctocephalus
  • South American fur seal (A. australis)
  • Australasian fur seal (A. forsteri)
  • Galápagos fur seal (A. galapagoensis)
  • Antarctic fur seal (A. gazella)
  • Juan Fernández fur seal (A. philippii)
  • Brown fur seal (A. pusillus)
  • Guadalupe fur seal (A. townsendi)
  • Subantarctic fur seal (A. tropicalis)
Callorhinus
  • Northern fur seal (C. ursinus)
Eumetopias
  • Steller sea lion (E. jubatus)
Neophoca
  • Australian sea lion (N. cinerea)
Otaria
  • South American sea lion (O. flavescens)
Phocarctos
  • New Zealand sea lion (P. hookeri)
Zalophus
  • California sea lion (Z. californianus)
  • Galápagos sea lion (Z. wollebaeki)
Odobenidae
(Pinniped inclusive)
Odobenus
  • Walrus (O. rosmarus)
Phocidae
(Earless seals)
(Pinniped inclusive)
Cystophora
  • Hooded seal (C. cristata)
Erignathus
  • Bearded seal (E. barbatus)
Halichoerus
  • Grey seal (H. grypus)
Histriophoca
  • Ribbon seal (H. fasciata)
Hydrurga
  • Leopard seal (H. leptonyx)
Leptonychotes
  • Weddell seal (L. weddellii)
Lobodon
  • Crabeater seal (L. carcinophagus)
Mirounga
(Elephant seals)
  • Northern elephant seal (M. angustirostris)
  • Southern elephant seal (M. leonina)
Monachus
  • Mediterranean monk seal (M. monachus)
Neomonachus
  • Hawaiian monk seal (N. schauinslandi)
Ommatophoca
  • Ross seal (O. rossi)
Pagophilus
  • Harp seal (P. groenlandicus)
Phoca
  • Spotted seal (P. largha)
  • Harbor seal (P. vitulina)
Pusa
  • Caspian seal (P. caspica)
  • Ringed seal (P. hispida)
  • Baikal seal (P. sibirica)
Canidae
Large family listed below
Mustelidae
Large family listed below
Family Canidae (includes dogs)
Atelocynus
  • Short-eared dog (A. microtis)
Canis
  • Golden jackal (C. aureus)
  • Domestic dog (C. familiaris)
  • Coyote (C. latrans)
  • African wolf (C. lupaster)
  • Wolf (C. lupus)
  • Eastern wolf (C. lycaon)
  • Red wolf (C. rufus)
  • Ethiopian wolf (C. simensis)
Cerdocyon
  • Crab-eating fox (C. thous)
Chrysocyon
  • Maned wolf (C. brachyurus)
Cuon
  • Dhole (C. alpinus)
Lupulella
  • Side-striped jackal (L. adustus)
  • Black-backed jackal (L. mesomelas)
Lycalopex
  • Culpeo (L. culpaeus)
  • Darwin's fox (L. fulvipes)
  • South American gray fox (L. griseus)
  • Pampas fox (L. gymnocercus)
  • Sechuran fox (L. sechurae)
  • Hoary fox (L. vetulus)
Lycaon
  • African wild dog (L. pictus)
Nyctereutes
  • Common raccoon dog (N. procyonoides)
  • Japanese raccoon dog (N. viverrinus)
Otocyon
  • Bat-eared fox (O. megalotis)
Speothos
  • Bush dog (S. venaticus)
Urocyon
  • Gray fox (U. cinereoargenteus)
  • Island fox (U. littoralis)
Vulpes
(Foxes)
  • Bengal fox (V. bengalensis)
  • Blanford's fox (V. cana)
  • Cape fox (V. chama)
  • Corsac fox (V. corsac)
  • Tibetan fox (V. ferrilata)
  • Arctic fox (V. lagopus)
  • Kit fox (V. macrotis)
  • Pale fox (V. pallida)
  • Rüppell's fox (V. rueppelli)
  • Swift fox (V. velox)
  • Red fox (V. vulpes)
  • Fennec fox (V. zerda)
Family Mustelidae
Helictidinae
(Ferret-badgers)
Melogale
  • Vietnam ferret-badger (M. cucphuongensis)
  • Bornean ferret badger (M. everetti)
  • Chinese ferret-badger (M. moschata)
  • Javan ferret-badger (M. orientalis)
  • Burmese ferret-badger (M. personata)
  • Formosan ferret-badger (M. subaurantiaca)
Guloninae
(Martens and wolverines)
Eira
  • Tayra (E. barbara)
Gulo
  • Wolverine (G. gulo)
Martes
(Martens)
  • American marten (M. americana)
  • Pacific marten (M. caurina)
  • Yellow-throated marten (M. flavigula)
  • Beech marten (M. foina)
  • Nilgiri marten (M. gwatkinsii)
  • European pine marten (M. martes)
  • Japanese marten (M. melampus)
  • Sable (M. zibellina)
Pekania
  • Fisher (P. pennanti)
Ictonychinae
(African polecats and grisons)
Galictis
  • Lesser grison (G. cuja)
  • Greater grison (G. vittata)
Ictonyx
  • Saharan striped polecat (I. libyca)
  • Striped polecat (I. striatus)
Lyncodon
  • Patagonian weasel (L. patagonicus)
Poecilogale
  • African striped weasel (P. albinucha)
Vormela
  • Marbled polecat (V. peregusna)
Lutrinae
(Otters)
Aonyx
  • African clawless otter (A. capensis)
  • Asian small-clawed otter (A. cinereus)
  • Congo clawless otter (A. congicus)
Enhydra
  • Sea otter (E. lutris)
Hydrictis
  • Spotted-necked otter (H. maculicollis)
Lontra
  • North American river otter (L. canadensis)
  • Marine otter (L. felina)
  • Neotropical otter (L. longicaudis)
  • Southern river otter (L. provocax)
Lutra
  • Eurasian otter (L. lutra)
  • Hairy-nosed otter (L. sumatrana)
Lutrogale
  • Smooth-coated otter (L. perspicillata)
Pteronura
  • Giant otter (P. brasiliensis)
Melinae
(Eurasian badgers)
Arctonyx
  • Northern hog badger (A. albogularis)
  • Greater hog badger (A. collaris)
  • Sumatran hog badger (A. hoevenii)
Meles
  • Japanese badger (M. anakuma)
  • Caucasian badger (M. canescens)
  • Asian badger (M. leucurus)
  • European badger (M. meles)
Mellivorinae
Mellivora
  • Honey badger (M. capensis)
Mustelinae
(Weasels and minks)
Mustela
(Weasels and ferrets)
  • Sichuan weasel (M. aistoodonnivalis)
  • Mountain weasel (M. altaica)
  • Stoat/Beringian ermine (M. erminea)
  • Steppe polecat (M. eversmannii)
  • Ferret (M. furo)
  • Haida ermine (M. haidarum)
  • Japanese weasel (M. itatsi)
  • Yellow-bellied weasel (M. kathiah)
  • European mink (M. lutreola)
  • Indonesian mountain weasel (M. lutreolina)
  • Black-footed ferret (M. nigripes)
  • Least weasel (M. nivalis)
  • Malayan weasel (M. nudipes)
  • European polecat (M. putorius)
  • American ermine (M. richardsonii)
  • Siberian weasel (M. sibirica)
  • Back-striped weasel (M. strigidorsa)
Neogale
  • Amazon weasel (N. africana)
  • Colombian weasel (N. felipei)
  • Long-tailed weasel (N. frenata)
  • American mink (N. vison)
Taxidiinae
Taxidea
  • American badger (T. taxus)
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Genera of civets, mongooses, hyenas, cats, and their extinct allies
Palaeogalidae
Nimravidae
Aeluroidea
Nandiniidae
Viverroidea
Viverridae
Genettinae
Hemigalinae
Paradoxurinae
Viverrinae
Herpestoidea
Herpestidae
    • See below↓
Hyaenidae
    • See below↓
Feloidea
    • See below↓
Eupleridae
Euplerinae
Galidiinae
Herpestidae
  • Kichechia
  • Legetetia
  • Leptoplesictis
  • Ugandictis
Herpestinae
Mungotinae
Percrocutidae?
Lophocyonidae
Hyaenidae
  • Allohyaena
  • Belbus
  • Metahyaena
  • Pliocrocuta
  • Tongxinictis
  • Werdelinus
Ictitheriinae
Protelinae
Hyaeninae
Barbourofelidae
  • (See Barbourofelidae)
Prionodontidae
Felidae
Proailurinae
Felinae
Machairodontinae
Pantherinae
Taxon identifiers
Fossa fossana