Pacific sleeper shark

Species of shark

Pacific sleeper shark
Conservation status

Near Threatened  (IUCN 3.1)[1]
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Chondrichthyes
Subclass: Elasmobranchii
Subdivision: Selachimorpha
Order: Squaliformes
Family: Somniosidae
Genus: Somniosus
Species:
S. pacificus
Binomial name
Somniosus pacificus
Bigelow & Schroeder, 1944
Range of the Pacific sleeper shark
Pacific sleeper shark carcasses

The Pacific sleeper shark (Somniosus pacificus) is a sleeper shark of the family Somniosidae, found in the North Pacific on continental shelves and slopes in Arctic and temperate waters between latitudes 70°N and 22°N and in at least two places in the western tropical Pacific near Palau and the Solomon Islands, from the surface to 2,000 metres (6,600 ft) deep.[1][2] The first evidence of the sharks in the western tropical Pacific emerged from a National Geographic video taken near the Solomon Islands in 2015.[3] Its length is up to 4.4 m (14 ft), although it could possibly reach lengths in excess of 7 m (23 ft).[2]

Feeding habits

Pacific sleeper sharks, which are thought to be both predators and scavengers, can glide through the water with little body movement and little hydrodynamic noise, making them successful stealth predators. They feed by means of suction and cutting of their prey. They have large mouths that can essentially inhale prey and their teeth cut up any pieces that are too large to swallow. They show a characteristic rolling motion of the head when feeding. Only in Alaska has the shark's diet been studied - most sharks' stomachs contain remains of giant Pacific octopus. They are also known to feed on bottom-dwelling teleost fishes, as well as soles, flounders, Alaska pollock, rockfishes, shrimps, hermit crabs, and even marine snails. Larger Pacific sleeper sharks are also found to feed on fast-swimming prey such as squids, Pacific salmon, and harbor porpoises. The diet of the Pacific sleeper shark seems to broaden as they increase in size. For example, a 3.7-m female shark found off Trinidad, California was found to have fed mostly on giant squid. Sleeper sharks found in Alaskan waters from 2 to 3 m (6.6 to 9.8 ft) seem to feed mostly on flounder, pollock, and cephalopods, while sleeper sharks 3.3 to 4.25 m (10.8 to 13.9 ft) long seem to consume teleosts and cephalopods, as well as marine mammals. A recent study in the Gulf of Alaska suggests that sleeper sharks may prey on juvenile Steller sea lions.[4]

Reproduction

Very little is known about the early life of Pacific sleeper sharks. They are believed to produce eggs that hatch inside the female's body (reproduction is ovoviviparous), but gestation time is unknown and litter sizes are thought to be about 10 pups. Its length at birth is about 42 cm (1.38 ft) or less.[5]

Size

The average mature size is 3.65 m (12.0 ft) and 318–363 kg (701–800 lb). The largest Pacific sleeper shark verified in size measured 4.4 m (14 ft) long and weighed 888 kg (1,958 lb), although it could possibly reach 7 m (23 ft) or more.[2][6] In 1989, an enormous Pacific sleeper shark was attracted to a bait in deep water outside Tokyo Bay, Japan, and filmed. The shark was estimated by Eugenie Clark to be about 7 m (23 ft) long.[5] A single unconfirmed account exists of an enormous Pacific sleeper shark that potentially measured more than 9.2 m (30 ft) long.[7] If true, this would make the species the largest extant macro-predatory shark, and the third largest shark overall after the whale shark and the basking shark.

Adaptations

Due to living in frigid depths, the sleeper shark's liver oil does not contain squalene, which would solidify into a dense, nonbuoyant mass. Instead, the low-density compounds in the sharks' liver are diacylglyceryl ethers and triacylglycerol, which maintain their fluidity even at the lowest temperatures. Also, they store very little urea in their skin (like many deep sea sharks), but like other elasmobranchs, have high concentrations of urea and trimethylamine oxide (nitrogenous waste products) in their tissues as osmoprotectants and to increase their buoyancy.[8] Trimethylamine oxide also serves to counteract the protein-destabilizing tendencies of urea[9] and pressure.[10] Its presence in the tissues of both elasmobranch and teleost fish has been found to increase with depth.[10][11]

Because food is relatively scarce on the deep sea floor, the sleeper shark is able to store food in its capacious stomach. The sleeper shark's jaws are able to produce a powerful bite due to their short and transverse shape. The upper jaw teeth of the sleeper shark are spike-like, while the lower jaw teeth are oblique cusps and overlapping bases. This arrangement allows grasping and sawing of food too large to swallow. Pacific sleeper sharks have a short caudal fin, which allows them to store energy for fast and violent bursts of energy to catch prey.[5]

In 2015 a pacific sleeper shark was filmed near the Solomon Islands underneath an active volcano. [12][13]

Known predators

Sleeper sharks are preyed on by the offshore ecotype of killer whale off British Columbia.[14] In addition, like the Greenland shark, the parasitic copepod Ommatokoita elongata can often be observed consuming the shark's corneal tissue.

See also

  • iconSharks portal

References

  1. ^ a b Rigby, C.L.; Derrick, D.; Dyldin, Y.V.; Ebert, D.A.; Herman, K.; Ho, H.; Hsu, H.; Ishihara, H.; Jeong, C.-H.; Pacoureau, N.; Semba, Y.; Tanaka, S.; Volvenko, I.V.; Walls, R.H.L.; Yamaguchi, A. (2021). "Somniosus pacificus". IUCN Red List of Threatened Species. 2021: e.T161403A887942. doi:10.2305/IUCN.UK.2021-2.RLTS.T161403A887942.en. Retrieved 19 November 2021.
  2. ^ a b c Froese, Rainer; Pauly, Daniel (eds.) (2016). "Somniosus pacificus" in FishBase. March 2016 version.
  3. ^ Claassens, L., Phillips, B., Ebert, D. A., Delaney, D., Henning, B., Nestor, V., Ililau, A., & Giddens, J. (2023). First records of the Pacific sleeper shark Somniosus cf. pacificus in the western tropical Pacific. Journal of fish biology, 10.1111/jfb.15487. Advance online publication. https://doi.org/10.1111/jfb.15487
  4. ^ Markus Horning & Jo-Ann Mellish (2014). "In cold blood: evidence of Pacific sleeper shark (Somniosus pacificus) predation on Steller sea lions (Eumetopias jubatus) in the Gulf of Alaska" (PDF). Fishery Bulletin. 112 (4): 297–310. doi:10.7755/FB.112.4.6. Archived from the original on 2020-05-29. Retrieved 2018-10-24.
  5. ^ a b c Martin, R. Aidan. "Elasmo Research". ReefQuest. Archived from the original on 20 April 2013. Retrieved 6 May 2009.
  6. ^ Castro, José I., The Sharks of North America. Oxford University Press (2011), ISBN 978-0-19-539294-4
  7. ^ The New World of the Oceans: Men and Oceanography
  8. ^ Withers, P. C.; Morrison, G.; Guppy, M. (May 1994). "Buoyancy Role of Urea and TMAO in an Elasmobranch Fish, the Port Jackson Shark, Heterodontus portusjacksoni". Physiological Zoology. 67 (3): 693–705. doi:10.1086/physzool.67.3.30163765. JSTOR 30163765. S2CID 100989392.
  9. ^ Bennion, B. J.; Daggett, V. (27 April 2004). "Counteraction of urea-induced protein denaturation by trimethylamine N-oxide: a chemical chaperone at atomic resolution". Proceedings of the National Academy of Sciences. 101 (17): 6433–6438. Bibcode:2004PNAS..101.6433B. doi:10.1073/pnas.0308633101. PMC 404062. PMID 15096583.
  10. ^ a b Yancey, P. H.; Gerringer, M. E.; Drazen, J. C.; Rowden, A. A.; Jamieson, A. (2014-03-03). "Marine fish may be biochemically constrained from inhabiting the deepest ocean depths". Proceedings of the National Academy of Sciences. 111 (12): 4461–4465. Bibcode:2014PNAS..111.4461Y. doi:10.1073/pnas.1322003111. PMC 3970477. PMID 24591588.
  11. ^ Treberg, J. R.; Driedzic, W. R. (2002-05-30). "Elevated levels of trimethylamine oxide in deep-sea fish: evidence for synthesis and intertissue physiological importance". Journal of Experimental Zoology. 293 (1): 39–45. doi:10.1002/jez.10109. PMID 12115917.
  12. ^ Mansfield, Katie (20 April 2017). "Mysterious sharks living INSIDE active underwater VOLCANO investigated by robots". express.co.uk. Archived from the original on 22 March 2018. Retrieved 21 March 2018.
  13. ^ National Geographic (8 July 2015). "Rarely Seen Shark Filmed Near Underwater Volcano - National Geographic". Archived from the original on 14 April 2019. Retrieved 21 March 2018 – via YouTube.
  14. ^ Keven Drews; The Canadian Press (5 September 2011). "Killer whales feast on sharks off B.C. coast". The Canadian Press. Archived from the original on 13 August 2013. Retrieved 5 Sep 2011.
General references
  • "Somniosus pacificus". Integrated Taxonomic Information System. Retrieved 23 January 2006.
  • "New giant squid predator found". BBC News. 2004-01-08. Retrieved February 14, 2007.
  • Castro, Jose. "Pacific Sleeper Sharks (Somniosus pacificus)." Conservation Science Institute. 1983. [1].
  • Martin, R. A. "Pacific Sleeper Shark Bibliography." Biology of Sharks and Reys. ReefQuest Centre for Shark Research.[2].
  • "Megalodon caught on tape." My Paranormal Life. Google. [3]. (Erroneously labelled footage of a sleeper shark)
  • Carroll, Amy. "Sleeper Sharks: Awake and Hungry Sleeper sharks Not Culprits in Sea Lion Declines." Alaska Fish and Wildlife News. 1999. Alaska Department of Fish and Game.[4].

External links

  • Paul Rincon (January 8, 2004). "New giant squid predator found". BBC News.
  • ORCA Video of a sleeper shark
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Extant shark species
Order Carcharhiniformes (Ground sharks)
Hemigaleidae
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Scylliogaleus
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Family Carcharhinidae (Requiem sharks)
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Isogomphodon
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Lamiopsis
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Loxodon
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Nasolamia
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Negaprion
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Prionace
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Rhizoprionodon
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Scoliodon
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Triaenodon
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Family Scyliorhinidae (Catsharks)
Apristurus
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  • A. pinguis
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  • Pale catshark (A. sibogae)
  • South China catshark (A. sinensis)
  • Spongehead catshark (A. spongiceps)
  • Panama ghost catshark (A. stenseni)
Asymbolus
  • Australian spotted catshark (A. analis)
  • A. funebris
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  • A. parvus
  • A. rubiginosus
  • Variegated catshark (A. submaculatus)
  • Gulf catshark (A. vincenti)
Atelomycterus
  • A. baliensis
  • Banded sand catshark (A. fasciatus)
  • Australian marbled catshark (A. macleayi)
  • Coral catshark (A. marmoratus)
Aulohalaelurus
  • New Caledonia catshark (A. kanakorum)
  • Australian blackspotted catshark (A. labiosus)
Cephaloscyllium
  • Whitefin swellshark (C. albipinnum)
  • Circle-blotch pygmy swellshark (C. circulopullum)
  • Cook's swellshark (C. cooki)
  • Reticulated swellshark (C. fasciatum)
  • Formosa swellshark (C. formosanum)
  • Australian reticulate swellshark (C. hicosellum)
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  • Australian swellshark (C. laticeps)
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  • Leopard-spotted swellshark (C. pardelotum)
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  • Flagtail swellshark (C. signourum)
  • Indian swellshark (C. silasi)
  • Speckled swellshark (C. speccum)
  • Balloon shark (C. sufflans)
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  • Saddled swellshark (C. variegatum)
  • Swellshark (C. ventriosum)
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Cephalurus
  • Lollipop catshark (C. cephalus)
Figaro
  • Australian sawtail catshark (F. boardmani)
  • Northern sawtail catshark (F. striatus)
Galeus
  • Antilles catshark (G. antillensis)
  • Roughtail catshark (G. arae)
  • Atlantic sawtail cat shark (G. atlanticus)
  • Longfin sawtail cat shark (G. cadenati)
  • Gecko catshark (G. eastmani)
  • Slender sawtail catshark (G. gracilis)
  • Longnose sawtail cat shark (G. longirostris)
  • Blackmouth catshark (G. melastomus)
  • Southern sawtail catshark (G. mincaronei)
  • Mouse catshark (G. murinus)
  • Broadfin sawtail catshark (G. nipponensis)
  • Peppered catshark (G. piperatus)
  • African sawtail catshark (G. polli)
  • G. priapus
  • Blacktip sawtail catshark (G. sauteri)
  • Dwarf sawtail catshark (G. schultzi)
  • Springer's sawtail cat shark (G. springeri)
Halaelurus
  • Arabian catshark (H. alcockii)
  • Speckled catshark (H. boesemani)
  • Blackspotted catshark (H. buergeri)
  • Dusky catshark (H. canescens)
  • Broadhead cat shark (H. clevai)
  • New Zealand catshark (H. dawsoni)
  • Bristly catshark (H. hispidus)
  • Spotless catshark (H. immaculatus)
  • Lined catshark (H. lineatus)
  • Mud catshark (H. lutarius)
  • Tiger catshark (H. natalensis)
  • Quagga catshark (H. quagga)
Haploblepharus
  • Puffadder shyshark (H. edwardsii)
  • Brown shyshark (H. fuscus)
  • Natal shyshark (H. kistnasamyi)
  • Dark shyshark (H. pictus)
Holohalaelurus
  • H. favus
  • H. grennian
  • Crying izak (H. melanostigma)
  • African spotted catshark (H. punctatus)
  • Izak catshark (H. regani)
Parmaturus
  • White-tip catshark (P. albimarginatus)
  • White-clasper catshark (P. albipenis)
  • Beige catshark (P. bigus)
  • Campeche catshark (P. campechiensis)
  • Velvet catshark (P. lanatus)
  • McMillan's catshark (P. macmillani)
  • Blackgill catshark (P. melanobranchus)
  • Salamander shark (P. pilosus)
  • Filetail catshark (P. xaniurus)
  • Shorttail catshark (Parmaturus sp. A)
Pentanchus
  • Onefin catshark (P. profundicolus)
Poroderma
  • Pyjama catshark (P. africanum)
  • Leopard catshark (P. pantherinum)
Schroederichthys
  • Narrowmouthed catshark (S. bivius)
  • Redspotted catshark (S. chilensis)
  • Narrowtail catshark (S. maculatus)
  • Lizard catshark (S. saurisqualus)
  • Slender catshark (S. tenuis)
Scyliorhinus
  • Polkadot catshark (S. besnardi)
  • Boa catshark (S. boa)
  • Small-spotted catshark (S. canicula)
  • Yellowspotted catshark (S. capensis)
  • West African catshark (S. cervigoni)
  • Comoro cat shark (S. comoroensis)
  • Brownspotted catshark (S. garmani)
  • Freckled catshark (S. haeckelii)
  • Whitesaddled catshark (S. hesperius)
  • Blotched catshark (S. meadi)
  • Chain catshark (S. retifer)
  • Nursehound (S. stellaris)
  • Izu cat shark (S. tokubee)
  • Cloudy catshark (S. torazame)
  • Dwarf catshark (S. torrei)
Order Echinorhiniformes (Bramble sharks)
Echinorhinidae
Echinorhinus
  • Bramble shark (E. brucus)
  • Prickly shark (E. cookei)
Order Heterodontiformes (Bullhead sharks)
Heterodontidae
Heterodontus
  • Horn shark (H. francisci)
  • Crested bullhead shark (H. galeatus)
  • Japanese bullhead shark (H. japonicus)
  • Mexican hornshark (H. mexicanus)
  • Oman bullhead shark (H. omanensis)
  • Port Jackson shark (H. portusjacksoni)
  • Galapagos bullhead shark (H. quoyi)
  • Whitespotted bullhead shark (H. ramalheira)
  • Zebra bullhead shark (H. zebra)
Chlamydoselachidae
Chlamydoselachus
  • Frilled shark (C. anguineus)
  • Southern African frilled shark (C. africana)
Hexanchidae
(Cow sharks)
Heptranchias
  • Sharpnose sevengill shark (H. perlo)
Hexanchus
  • Bluntnose sixgill shark (H. griseus)
  • Bigeyed sixgill shark (H. nakamurai)
Notorynchus
  • Broadnose sevengill shark (N. cepedianus)
Order Lamniformes (Mackerel sharks)
Alopiidae
Alopias
(Thresher sharks)
  • Pelagic thresher (A. pelagicus)
  • Bigeye thresher (A. superciliosus)
  • Common thresher (A. vulpinus)
  • Alopias sp. (A. sp.)
Cetorhinidae
Cetorhinus
  • Basking shark (C. maximus)
Lamnidae
Carcharodon
  • Great white shark (C. carcharias)
Isurus
  • Shortfin mako shark (I. oxyrinchus)
  • Longfin mako shark (I. paucus)
Lamna
  • Salmon shark (L. ditropis)
  • Porbeagle (L. nasus)
Megachasmidae
Megachasma
  • Megamouth shark (M. pelagios)
Mitsukurinidae
Mitsukurina
  • Goblin shark (M. owstoni)
Odontaspididae
Carcharias
  • Grey nurse shark (C. taurus)
  • Indian sand tiger (C. tricuspidatus)
Odontaspis
  • Smalltooth sand tiger (O. ferox)
  • Bigeye sand tiger (O. noronhai)
Pseudocarchariidae
Pseudocarcharias
  • Crocodile shark (P. kamoharai)
Order Orectolobiformes (Carpet sharks)
Brachaeluridae
Brachaelurus
  • Blind shark (B. waddi)
Heteroscyllium
  • Bluegrey carpetshark (H. colcloughi)
Ginglymostomatidae
(Nurse sharks)
Ginglymostoma
  • Nurse shark (G. cirratum)
Nebrius
  • Tawny nurse shark (N. ferrugineus)
Pseudoginglymostoma
  • Short-tail nurse shark (P. brevicaudatum)
Hemiscylliidae
(Bamboo sharks)
Chiloscyllium
  • Arabian carpetshark (C. arabicum)
  • Burmese bamboo shark (C. burmensis)
  • Bluespotted bamboo shark (C. caerulopunctatum)
  • Grey bamboo shark (C. griseum)
  • Hasselt's bamboo shark (C. hasseltii)
  • Slender bamboo shark (C. indicum)
  • Whitespotted bamboo shark (C. plagiosum)
  • Brownbanded bamboo shark (C. punctatum)
Hemiscyllium
  • Indonesian speckled carpetshark (H. freycineti)
  • H. galei
  • Papuan epaulette shark (H. hallstromi)
  • H. henryi
  • Epaulette shark (H. ocellatum)
  • Hooded carpetshark (H. strahani)
  • Speckled carpetshark (H. trispeculare)
Orectolobidae
(Wobbegongs)
Eucrossorhinus
  • Tasselled wobbegong (E. dasypogon)
Orectolobus
  • Floral banded wobbegong (O. floridus)
  • Banded wobbegong (O. halei)
  • Western wobbegong (O. hutchinsi)
  • Japanese wobbegong (O. japonicus)
  • Spotted wobbegong (O. maculatus)
  • Ornate wobbegong (O. ornatus)
  • Dwarf spotted wobbegong (O. parvimaculatus)
  • Network wobbegong (O. reticulatus)
  • Northern wobbegong (O. wardi)
Sutorectus
  • Cobbler wobbegong (S. tentaculatus)
Parascylliidae
(Collared carpet sharks)
Cirrhoscyllium
  • Barbelthroat carpetshark (C. expolitum)
  • Taiwan saddled carpetshark (C. formosanum)
  • Saddle carpetshark (C. japonicum)
Parascyllium
  • Collared carpetshark (P. collare)
  • Rusty carpetshark (P. ferrugineum)
  • Ginger carpetshark (P. sparsimaculatum)
  • Necklace carpetshark (P. variolatum)
Rhincodontidae
Rhincodon
  • Whale shark (R. typus)
Stegostomatidae
Stegostoma
  • Zebra shark (S. fasciatum)
Order Pristiophoriformes (Sawsharks)
Pristiophoridae
Pliotrema
  • Sixgill sawshark (P. warreni)
Pristiophorus
  • Longnose sawshark (P. cirratus)
  • Tropical sawshark (P. delicatus)
  • Japanese sawshark (P. japonicus)
  • Shortnose sawshark (P. nudipinnis)
  • Bahamas sawshark (P. schroederi)
  • Eastern Australian sawshark (Pristiophorus peroniensis)
  • Philippine sawshark (Pristiophorus sp. C)
  • Dwarf sawshark (Pristiophorus sp. D)
Centrophoridae
(Gulper sharks)
Centrophorus
  • Needle dogfish (C. acus)
  • Dwarf gulper shark (C. atromarginatus)
  • Gulper shark (C. granulosus)
  • Dumb gulper shark (C. harrissoni)
  • Blackfin gulper shark (C. isodon)
  • Lowfin gulper shark (C. lusitanicus)
  • Smallfin gulper shark (C. moluccensis)
  • Taiwan gulper shark (C. niaukang)
  • Leafscale gulper shark (C. squamosus)
  • Mosaic gulper shark (C. tessellatus)
  • Little gulper shark (C. uyato)
Deania
  • Birdbeak dogfish (D. calcea)
  • Rough longnose dogfish (D. hystricosa)
  • Arrowhead dogfish (D. profundorum)
  • Longsnout dogfish (D. quadrispinosum)
Dalatiidae
Euprotomicroides
  • Taillight shark (E. zantedeschia)
Heteroscymnoides
  • Longnose pygmy shark (H. marleyi)
Mollisquama
  • Pocket shark (M. parini)
Dalatias
  • Kitefin shark (D. licha)
Isistius
  • Cookiecutter shark (I. brasiliensis)
  • South China cookiecutter shark (I. labialis)
  • Largetooth cookiecutter shark (I. plutodus)
Euprotomicrus
  • Pygmy shark (E. bispinatus)
Squaliolus
  • Smalleye pygmy shark (S. aliae)
  • Spined pygmy shark (S. laticaudus)
Etmopteridae
Aculeola
  • Hooktooth dogfish (A. nigra)
Centroscyllium
  • Highfin dogfish (C. excelsum)
  • Black dogfish (C. fabricii)
  • Granular dogfish (C. granulatum)
  • Bareskin dogfish (C. kamoharai)
  • Combtooth dogfish (C. nigrum)
  • Ornate dogfish (C. ornatum)
  • Whitefin dogfish (C. ritteri)
Etmopterus
(Lantern sharks)
  • New Zealand lanternshark (E. baxteri)
  • Blurred lanternshark (E. bigelowi)
  • Shorttail lanternshark (E. brachyurus)
  • Lined lanternshark (E. bullisi)
  • E. burgessi
  • Cylindrical lanternshark (E. carteri)
  • Tailspot lanternshark (E. caudistigmus)
  • Combtooth lanternshark (E. decacuspidatus)
  • Pink lanternshark (E. dianthus)
  • E. dislineatus
  • Blackmouth lanternshark (E. evansi)
  • Pygmy lanternshark (E. fusus)
  • Broadbanded lanternshark (E. gracilispinis)
  • Southern lanternshark (E. granulosus)
  • Caribbean lanternshark (E. hillianus)
  • Smalleye lantern shark (E. litvinovi)
  • Blackbelly lanternshark (E. lucifer)
  • Slendertail lanternshark (E. molleri)
  • Dwarf lanternshark (E. perryi)
  • African lanternshark (E. polli)
  • Great lanternshark (E. princeps)
  • False lanternshark (E. pseudosqualiolus)
  • Smooth lanternshark (E. pusillus)
  • Dense-scale lantern shark (E. pycnolepis)
  • West Indian lanternshark (E. robinsi)
  • Fringefin lanternshark (E. schultzi)
  • Thorny lanternshark (E. sentosus)
  • Velvet belly lantern shark (E. spinax)
  • Splendid lanternshark (E. splendidus)
  • Tasmanian lanternshark (E. tasmaniensis)
  • Brown lanternshark (E. unicolor)
  • Hawaiian lanternshark (E. villosus)
  • Green lanternshark (E. virens)
Miroscyllium
  • Rasptooth dogfish (M. sheikoi)
Trigonognathus
  • Viper dogfish (T. kabeyai)
Oxynotidae
(Rough sharks)
Oxynotus
  • Prickly dogfish (O. bruniensis)
  • Caribbean roughshark (O. caribbaeus)
  • Angular roughshark (O. centrina)
  • Japanese roughshark (O. japonicus)
  • Sailfin roughshark (O. paradoxus)
Somniosidae
(Sleeper sharks)
Centroscymnus
  • Portuguese dogfish (C. coelolepis)
  • Shortnose velvet dogfish (C. cryptacanthus)
  • Roughskin dogfish (C. owstoni)
Centroselachus
  • Longnose velvet dogfish (C. crepidater)
Proscymnodon
  • Largespine velvet dogfish (P. macracanthus)
  • Plunket shark (P. plunketi)
Scymnodalatias
  • Whitetail dogfish (S. albicauda)
  • Azores dogfish (S. garricki)
  • Sparsetooth dogfish (S. oligodon)
  • Sherwood dogfish (S. sherwoodi)
Scymnodon
  • Smallmouth velvet dogfish (S. obscurus)
  • Knifetooth dogfish (S. ringens)
Somniosus
  • Southern sleeper shark (S. antarcticus)
  • Frog shark (S. longus )
  • Greenland shark (S. microcephalus)
  • Pacific sleeper shark (S. pacificus)
  • Little sleeper shark (S. rostratus)
Zameus
  • Japanese velvet dogfish (Z. ichiharai)
  • Velvet dogfish (Z. squamulosus)
Squalidae
(Dogfish sharks)
Cirrhigaleus
  • Roughskin spurdog (C. asper)
  • Mandarin dogfish (C. barbifer)
Squalus
(Spurdogs)
  • Spiny dogfish (S. acanthias)
  • Eastern highfin spurdog (S. albifrons)
  • S. acutirostris
  • Western highfin spurdog (S. altipinnis)
  • Longnose spurdog (S. blainville)
  • Fatspine spurdog (S. crassispinus)
  • Cuban dogfish (S. cubensis)
  • Eastern longnose spurdog (S. grahami)
  • Japanese spurdog (S. japonicus)
  • Shortnose spurdog (S. megalops)
  • Blacktailed spurdog (S. melanurus)
  • Shortspine spurdog (S. mitsukurii)
  • Bartail spurdog (S. notocaudatus)
  • Western longnose spurdog (S. nasutus)
  • Cyrano spurdog (S. rancureli)
  • Pacific spiny dogfish (S. suckleyi)
Order Squatiniformes (Angel sharks)
Squatinidae
Squatina
  • Sawback angelshark (S. aculeata)
  • African angelshark (S. africana)
  • Eastern Australian angelshark (Squatina albipunctata)
  • Argentine angelshark (S. argentina)
  • Chilean angelshark (S. armata)
  • Australian angelshark (S. australis)
  • Pacific angelshark (S. californica)
  • Sand devil (S. dumeril)
  • Taiwan angelshark (S. formosa)
  • Angular angel shark (S. guggenheim)
  • S. heteroptera
  • Japanese angelshark (S. japonica)
  • Indonesian angelshark (S. legnota)
  • Cortez angelshark (S. mexicana)
  • Clouded angelshark (S. nebulosa)
  • Smoothback angelshark (S. oculata)
  • S. punctata
  • Western Australian angelshark (Squatina pseudocellata)
  • Angelshark (S. squatina)
  • Ornate angelshark (S. tergocellata)
  • Ocellated angelshark (S. tergocellatoides)
Taxon identifiers
Somniosus pacificus
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