Phrynops

Genus of turtles

Phrynops

Phrynops hilarii
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Pleurodira
Family:	Chelidae
Subfamily:	Chelinae
Genus:	Phrynops Wagler 1830^[1]
Synonyms^[2]
Spatulemys Gray 1872^[3] Parahydraspis Wieland 1923^[4]

Sometimes called the bearded toadheads but better known by their scientific name of Phrynops this genus of turtles has often been a bit of a dumping ground for the short-necked South American turtles of the family Chelidae.

The genus is widely distributed in South America found in the Orinoco to Amazon and São Francisco to Paraná and adjacent river basins of Colombia, Venezuela, the Guianas, Brazil, Paraguay, Uruguay and northeastern Argentina (Iverson, 1992).^[5]

A turtle of the genus Phrynops generally achieves thermoregulation through areal basking. In order to best attain its goal, it tends to increase its basking time during mid-day during the winters to make up for the cold. Other factors such as food intake and reproduction underwater also influence the way it controls the temperature of its body. Although this is the main way it is able to maintain body temperature, food intake and reproduction rates are also influential in the matter.

Species

Listed alphabetically by specific name.^[6]

Phrynops geoffroanus (Schweigger, 1812) – Geoffroy's toadhead turtle, Geoffroy's side-necked turtle^[7]
Phrynops hilarii (A.M.C. Duméril & Bibron, 1835) – Hilaire's toadhead turtle, Hilaire's side-necked turtle^[8]
Phrynops tuberosus (W. Peters, 1870) – Cotinga River toadhead turtle^[9]
Phrynops williamsi Rhodin & Mittermeier, 1983 – William's toadhead turtle, William's South American side-necked turtle^[10]
†Phrynops paranaensis (Wieland, 1923)^[4]^[11]

References

^ Wagler, J. (1830). Natürliches System der Amphibien, mit vorangehender Classification der Säugthiere und Vögel. Ein Beitrag zur vergleichenden Zoologie. Munich, Stuttgart, and Tübingen: J.G. Cotta. vi + 354 pp. + 9 plates. (in German and Latin).
^ Turtle Taxonomy Working Group (van Dijk, P.P.; Iverson, J.B.; Rhodin, A.G.J.; Shaffer, H.B.; Bour, R.) (2014). Turtles of the World, 7th edition: annotated checklist of taxonomy, synonymy, distribution with maps, and conservation status". In: Rhodin, A.G.J.; Pritchard, P.C.H.; van Dijk, P.P.; Saumure, R.A.; Buhlmann, K.A.; Iverson, J.B.; Mittermeier, R.A. (Editors) (2014). "Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group". Chelonian Research Monographs 5 (7): 000.329–479, doi:10.3854/ crm.5.000.checklist.v7.2014.
^ Gray, J.E. (1872). "On Spatulemys Lasalæ, a new genus of Hydraspidæ from Rio Parana, Corrientes". Annals and Magazine of Natural History, Fourth Series 10: 463.
^ ^a ^b Wieland, George R. (1923). "A new Paraná pleurodiran". American Journal of Science, Series 5, 5: 1–14.
^ lverson, J.B. (1992). Revised Checklist with Distribution Maps of the Turtles of the World. Richmond, Indiana: Privately Printed. 363 pp.
^ Phrynops, The Reptile Database
^ Schweigger, Augustus F. (1812). "Prodromus monographiae Cheloniorum ". Königsberger Archiv für Naturwissenschaft und Mathematik 1: 271–368, 406–462. (Emys geoffroana, new species, pp. 301–302). (in Latin).
^ Duméril, André Marie Constant; Bibron, Gabriel (1835). Erpétologie Générale ou Histoire Naturelle Complète des Reptiles. Tome Second. Paris: Roret. 680 pp. (Platemys hilarii, new species, pp. 428–430). (in French).
^ Peters, Wilhem Carl Hartwig (1870). "Über Platemys tuberosa, eine neue Art von Schildkröte aus British-Guiana ". Monatsberichte der Königliche Preussischen Akademie der Wissenschaften zu Berlin. 1870 (Mai): 311-313. (Platemys tuberosa, new species, pp. 311–313 + Plates 1–2). (in German).
^ Rhodin, Anders G.J.; Mittermeier, Russell A. (1983). "Description of Phrynops williamsi, a new species of chelid turtle of the South American P. geoffroanus complex". pp. 58-73. In: Rhodin, A.G.J.; Miyata, K. (Editors) (1983). Advances in Herpetology and Evolutionary Biology. Essays in Honor of Ernest E. Williams. Cambridge, Massachusetts: Museum of Comparative Zoology.
^ Phrynops at Fossilworks.org

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Suborder

Superfamily

Family

Genus

Cryptodira

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

Side-necked turtle species

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines
Suborder: Pleurodira

Family

Acanthochelys

	Acanthochelys macrocephala Acanthochelys pallidipectoris Acanthochelys radiolata Acanthochelys spixii

Chelus

	Chelus fimbriata

Mesoclemmys

	Mesoclemmys dahli Mesoclemmys gibba Mesoclemmys jurutiensis Mesoclemmys nasuta Mesoclemmys perplexa Mesoclemmys raniceps Mesoclemmys sabiniparaensis Mesoclemmys tuberculata Mesoclemmys vanderhaegei †Mesoclemmys vanegasorum Mesoclemmys zuliae

Phrynops

	Phrynops geoffroanus Phrynops hilarii †Phrynops paranaensis Phrynops tuberosus Phrynops williamsi

Platemys

	Platemys platycephala

Ranacephala

	Ranacephala hogei

Rhinemys

	Rhinemys rufipes

Chelodininae

Chelodina

Chelodina	Chelodina canni Chelodina gunaleni Chelodina longicollis Chelodina mccordi †Chelodina murrayi Chelodina novaeguineae Chelodina pritchardi Chelodina reimanni Chelodina steindachneri
Chelydera	†Chelodina alanrixi Chelodina burrungandjii Chelodina expansa Chelodina insculpta Chelodina kuchlingi Chelodina kurrichalpongo Chelodina parkeri Chelodina rugosa Chelodina walloyarrina
Macrochelodina	Chelodina oblonga

Elseya

Elseya	Elseya dentata Elseya branderhorsti Elseya flaviventralis
Pelocomastes	Elseya albagula Elseya irwini Elseya lavarackorum †Elseya nadibajagu †Elseya uberrima
Hanwarachelys	Elseya caelatus Elseya novaeguineae Elseya orestiad Elseya rhodini Elseya schultzei

Elusor

	Elusor macrurus

Emydura

	Emydura australis Emydura gunaleni Emydura macquarii Emydura subglobosa Emydura tanybaraga Emydura victoriae

Myuchelys

	Myuchelys belli Myuchelys georgesi Myuchelys latisternum Myuchelys purvisi

Rheodytes

	Rheodytes leukops †Rheodytes devisi

Hydromedusinae

Hydromedusa

	Hydromedusa maximiliani Hydromedusa tectifera

Pseudemydurinae

Pseudemydura

	Pseudemydura umbrina

Pelomedusidae

Pelomedusa

	Pelomedusa subrufa

Pelusios

	Pelusios adansonii Pelusios bechuanicus Pelusios broadleyi Pelusios carinatus Pelusios castaneus Pelusios castanoides Pelusios chapini Pelusios cupulatta Pelusios gabonensis Pelusios marani Pelusios nanus Pelusios niger Pelusios rhodesianus Pelusios sinuatus Pelusios subniger Pelusios upembae Pelusios williamsi

Podocnemididae

Erymnochelys

	Erymnochelys madagascariensis

Peltocephalus

	Peltocephalus dumerilianus

Podocnemis

	Podocnemis erythrocephala Podocnemis expansa Podocnemis lewyana Podocnemis sextuberculata Podocnemis unifilis Podocnemis vogli

Phylogenetic arrangement of turtles based on Turtles of the World 2021 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. Key: †=extinct.

Taxon identifiers
Phrynops	Wikidata: Q2700403 Wikispecies: Phrynops ADW: Phrynops BOLD: 171545 CoL: 63ML5 EoL: 17992 Fossilworks: 37616 GBIF: 2442114 iNaturalist: 39617 IRMNG: 1287776 ITIS: 202123 NCBI: 8462 Open Tree of Life: 66456 WoRMS: 1056589