Sphaeroceroidea

Superfamily of flies

Lesser dung flies and allies
Limosina silvatica
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Section: Schizophora
Subsection: Acalyptratae
Superfamily: Sphaeroceroidea
Families
  • Chyromyidae
  • Heleomyzidae - incl. Trixoscelididae & Rhinotoridae
  • Heteromyzidae
  • Nannodastiidae
  • Sphaeroceridae

Sphaeroceroidea is a superfamily of flies. It includes the cosmopolitan families of Sphaeroceridae (small dung flies), Heleomyzidae, and Chyromyidae, as well as a few smaller groups.[1] It has about 2,600 species.[2]

Description

Sphaeroceroids tend to be small flies a few millimetres in length.[3][4][5] Each of the families has its own distinguishing traits. For example, Sphaeroceridae have black wings with an interrupted costa, and the first tarsomere of the hind leg is short and thick.[5] Nannodastiidae are even smaller than usual for sphaeroceroids, with adults being 0.70-1.25 mm long, and their legs lack long setae.[6]

Ecology

Larvae generally live and feed on various kinds of decaying organic matter, including manure, seaweed, fungi, rotting wood, compost and carrion.[3][4][5]

Nannodastiidae have a more restricted lifestyle. Adults have usually been found in tropical and subtropical beaches, often in caves or under cliff overhangs. The larvae are unknown, but they may be in the droppings of birds and mammals which are abundant in such habitats.[6]

Taxonomy

The taxonomic composition of Sphaeroceroidea has been disputed. In 2007, there was a proposal to combine the families Heleomyzidae and Sphaeroceridae into one family, Heteromyzidae.[7] Other, more recent studies have kept these two families separate.[1][2]

Phylogeny

A 2021 study confirmed that the superfamily is monophyletic. Additionally, it found that Sphaeroceroidea is the sister group to all other Schizophora.[2]

References

  1. ^ a b Kahanpää, Jere (2014-09-19). "Checklist of the fly families Chyromyidae and Heleomyzidae (Diptera) of Finland". ZooKeys (441). Pensoft Publishers: 319–324. doi:10.3897/zookeys.441.7507. ISSN 1313-2970. PMC 4200468. PMID 25337028.
  2. ^ a b c Bayless, Keith M.; Trautwein, Michelle D.; Meusemann, Karen; Shin, Seunggwan; Petersen, Malte; Donath, Alexander; Podsiadlowski, Lars; Mayer, Christoph; Niehuis, Oliver; Peters, Ralph S.; Meier, Rudolf; Kutty, Sujatha Narayanan; Liu, Shanlin; Zhou, Xin; Misof, Bernhard (2021). "Beyond Drosophila: resolving the rapid radiation of schizophoran flies with phylotranscriptomics". BMC Biology. 19 (1): 23. doi:10.1186/s12915-020-00944-8. ISSN 1741-7007. PMC 7871583. PMID 33557827.
  3. ^ a b "Family Chyromyidae". bugguide.net. Retrieved 2022-10-24.
  4. ^ a b "Family Heleomyzidae". bugguide.net. Retrieved 2022-10-24.
  5. ^ a b c "Family Sphaeroceridae - Lesser Dung Flies". bugguide.net. Retrieved 2022-10-24.
  6. ^ a b Papp, L.; Mathis, W. N. (2001). "A review of the family Nannodastiidae (Diptera)". Proceedings of the Entomological Society of Washington. 103 (2): 337–348.
  7. ^ McAlpine, David K. (2007-08-29). "Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with reconsideration of the status of families Heleomyzidae and Sphaeroceridae, and descriptions of femoral gland-baskets". Records of the Australian Museum. 59 (3): 143–219. doi:10.3853/j.0067-1975.59.2007.1487. ISSN 0067-1975. S2CID 87660505.
  • Data related to Sphaeroceroidea at Wikispecies
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Extant Diptera families
Suborder Nematocera
Axymyiomorpha
Culicomorpha
Culicoidea
  • Dixidae (meniscus midges)
  • Corethrellidae (frog-biting midges)
  • Chaoboridae (phantom midges)
  • Culicidae (mosquitoes)
Chironomoidea
  • Thaumaleidae (solitary midges)
  • Simuliidae (black flies)
  • Ceratopogonidae (biting midges)
  • Chironomidae (non-biting midges)
Blephariceromorpha
  • Blephariceridae (net-winged midges)
  • Deuterophlebiidae (mountain midges)
  • Nymphomyiidae
Bibionomorpha
Bibionoidea
  • Bibionidae (march flies, lovebugs)
Anisopodoidea
  • Anisopodidae (wood gnats)
Sciaroidea
(fungus gnats)
Perissommatomorpha
Psychodomorpha
Scatopsoidea
Psychodoidea
  • Psychodidae (moth flies)
Ptychopteromorpha
  • Ptychopteridae (phantom crane flies)
  • Tanyderidae (primitive crane flies)
Tipulomorpha
Trichoceroidea
  • Trichoceridae (winter crane flies)
Tipuloidea
(crane flies)
  • Cylindrotomidae (long-bodied crane flies)
  • Limoniidae (limoniid crane flies)
  • Pediciidae (hairy-eyed craneflies)
  • Tipulidae (large crane flies)
Suborder Brachycera
Asilomorpha
Asiloidea
Empidoidea
Nemestrinoidea
  • Acroceridae (small-headed flies)
  • Nemestrinidae (tangle-veined flies)
Muscomorpha
Aschiza
Platypezoidea
  • Ironomyiidae (ironic flies)
  • Lonchopteridae (spear-winged flies)
  • Opetiidae (flat-footed flies)
  • Phoridae (scuttle flies, coffin flies, humpbacked flies)
  • Platypezidae (flat-footed flies)
Syrphoidea
  • Pipunculidae (big-headed flies)
  • Syrphidae (hoverflies)
Schizophora
Acalyptratae
Conopoidea
  • Conopidae (thick-headed flies)
Tephritoidea
  • Pallopteridae (flutter flies)
  • Piophilidae (cheese flies)
  • Platystomatidae (signal flies)
  • Pyrgotidae
  • Richardiidae
  • Tephritidae (peacock flies)
  • Ulidiidae (picture-winged flies)
Nerioidea
  • Cypselosomatidae
  • Micropezidae (stilt-legged flies)
  • Neriidae (cactus flies, banana stalk flies)
Diopsoidea
Sciomyzoidea
Sphaeroceroidea
Lauxanioidea
Opomyzoidea
Ephydroidea
  • Camillidae
  • Curtonotidae (quasimodo flies)
  • Diastatidae (bog flies)
  • Drosophilidae (vinegar and fruit flies)
  • Ephydridae (shore flies)
  • Mormotomyiidae (frightful hairy fly)
Carnoidea
Lonchaeoidea
Calyptratae
Muscoidea
  • Anthomyiidae (cabbage flies)
  • Fanniidae (little house flies)
  • Muscidae (house flies, stable flies)
  • Scathophagidae (dung flies)
Oestroidea
Hippoboscoidea
  • Glossinidae (tsetse flies)
  • Hippoboscidae (louse flies)
  • Nycteribiidae (bat flies)
  • Streblidae (bat flies)
Stratiomyomorpha
Stratiomyoidea
  • Pantophthalmidae (timber flies)
  • Stratiomyidae (soldier flies)
  • Xylomyidae (wood soldier flies)
Tabanomorpha
Rhagionoidea
Tabanoidea
Vermileonomorpha
Vermileonoidea
Xylophagomorpha
Xylophagoidea
  • Xylophagidae (awl flies)
Taxon identifiers
Sphaeroceroidea