Timarcha

Timarcha
Timarcha tenebricosa
Nomenklatur
Timarcha
Latreille, 1829
Populærnavn
Bladbiller
Klassifikasjon
RikeDyreriket
RekkeLeddyr
KlasseInsekter
Ordenbiller
Familiebladbiller
Økologi
Antall arter: over 100
Utbredelse: Palearktis
Inndelt i
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Timarcha er en slekt av biller i delgruppen (underfamilien) Chrysomelinae.

Utseende

Forholdsvis store og kraftige, brede og høyt hvelvede, halvblanke, svarte bladbiller, kan ha avdempet metallglans.

Levevis

Timarcha-artene er store og tunge biller med tykt skall, De har reduserte flygevinger, kan ikke fly, og er generelt trege. De kan dermed ikke lett unnslippe rovdyr, men forsvarer seg delvis med sitt tykke, harde skall og delvis ved å skille ut giftig blod om de blir alvorlig truet. Dette danner påfallende, røde dråper og har gitt billene det engelske navnet "bloody-nose beetles". Artene lever på bakken i tørre, varme områder, for eksempel maquis. Billene er hovedsklaig nattaktive.

Utbredelse

Slekten er utbredt i Nord-Afrika, Sør- og Sentral.Europa, Midtøsten og Asia. På grunn av billenes begrensede mobilitet har det utviklet seg svært mange lokale arter i denne slekten [1]. Dessuten finnes det noen få arter på vestkysten av Nord-Amerika.

Systematisk inndeling

Timarcha goettingensis

Det er sannsynlig at en del av de navngitte formene bare er fargevarianter [1].

Treliste
  • orden biller (Coleoptera)
    • underorden Polyphaga
      • overfamilie Chrysomeloidea Latreille, 1802 (=Phytophaga)
        • familien bladbiller (Chrysomelidae Latreille, 1802)
          • underfamilien Bladbiller, Chrysomelinae Latreille, 1802
            • stammen Timarchini
              • slekten Timarcha Latreille, 1829
                • Underslekten Americanotimarcha Jolivet, 1948 - noen ganger regnet som en egen slekt, to arter i vestlige Nord-Amerika
                  • Timarcha cerdo Stål, 1863
                    • Timarcha cerdo cerdo Stål, 1863
                    • Timarcha cerdo leechi Jolivet, 1948
                  • Timarcha intricata Haldeman, 1853
                    • Timarcha intricata intricata Haldeman, 1853
                    • Timarcha intricata vandykei Jolivet, 1948
                    • Timarcha intricata viridis Jolivet, 1948
                    • Timarcha intricata yvettae Jolivet, 1948
                • Underslekten Metallotimarcha Motschulsky, 1860
                  • Timarcha corinthia Fairmaire, 1873
                    • Timarcha corinthia albanica Apfelbeck, 1912
                    • Timarcha corinthia corinthia Fairmaire, 1873
                  • Timarcha gibba (Hagenbach, 1825)
                  • Timarcha hummelii Faldermann, 1837
                    • Timarcha hummelii humellii Faldermann, 1837
                    • Timarcha hummelii starcki Bechyné, 1953
                  • Timarcha metallica Laicharting, 1781
                • Underslekten Timarcha (i snever forstand)
                  • Timarcha apricaria Waltl, 1835
                  • Timarcha apuana Daccordi & Ruffo, 1990
                  • Timarcha aurichalcea Bechyné, 1948
                  • Timarcha balearica Gory, 1829
                  • Timarcha calceata Perez, 1865
                  • Timarcha chalcosoma Fairmaire, 1868
                  • Timarcha coarcticollis Fairmaire, 1873
                    • Timarcha coarcticollis coarcticollis Fairmaire, 1873
                    • Timarcha coarcticollis paulinoi Kraatz, 1879
                  • Timarcha erosa Fairmaire, 1873
                    • Timarcha erosa erosa Fairmaire, 1873
                    • Timarcha erosa vermiculata Fairmaire, 1880
                  • Timarcha espanoli Bechyné, 1948
                  • Timarcha fallax Perez, 1865
                    • Timarcha fallax fallax Perez, 1865
                    • Timarcha fallax piochardi Fairmaire, 1873
                    • Timarcha fallax sagrensis Kuntzen, 1911
                    • Timarcha fallax tortosensis Bechyné, 1948
                  • Timarcha granadensis Bechyné, 1948
                    • Timarcha granadensis dislocata Bechyné, 1948
                    • Timarcha granadensis granadensis Bechyné, 1948
                  • Timarcha gravis Rosenhauer, 1856
                  • Timarcha guipuzcoana Bechyné
                  • Timarcha heydeni Weise, 1882
                  • Timarcha hispanica Herrich-Schaeffer, 1838
                    • Timarcha hispanica hispanica Herrich-Schaeffer, 1838
                    • Timarcha hispanica laevisterna Fairmaire, 1880
                    • Timarcha hispanica rugipennis Perez, 1865
                    • Timarcha hispanica sobrina Fairmaire, 1884
                    • Timarcha hispanica transversicollis Fairmaire, 1880
                  • Timarcha insparsa Rosenhauer, 1856
                  • Timarcha intermedia Herrich-Schaeffer, 1838
                  • Timarcha kiesenwetteri Kraatz, 1879
                  • Timarcha marginicollis Rosenhauer, 1856
                  • Timarcha melica Bechyné, 1947
                  • Timarcha nicaeensis Villa, 1853
                    • Timarcha nicaeensis italica Herrich-Schäffer, 1838
                    • Timarcha nicaeensis nicaeensis Villa, 1853
                    • Timarcha nicaeensis sontiaca Müller, 1948
                  • Timarcha parvicollis Rosenhauer, 1856
                    • Timarcha parvicollis elocata Bechyné, 1953
                    • Timarcha parvicollis nitendula Bechyné, 1953
                    • Timarcha parvicollis parvicollis Rosenhauer, 1856
                  • Timarcha pimelioides Herrich-Schaeffer, 1838
                  • Timarcha rugulosa Herrich-Schaeffer, 1838
                    • Timarcha rugulosa globata Fairmaire, 1873
                    • Timarcha rugulosa lomnickii Miller, 1867
                    • Timarcha rugulosa rugulosa Herrich-Schaeffer, 1838
                    • Timarcha rugulosa sinaiana Bechyné, 1947
                    • Timarcha rugulosa stichai Bechyné, 1947
                  • Timarcha seidlitzi Kraatz, 1879
                  • Timarcha strophium Weise, 1888
                  • Timarcha tenebricosa (Fabricius, 1775)
                    • Timarcha tenebricosa anglica Bechyné, 1945
                    • Timarcha tenebricosa angusticollis Fairmaire, 1873
                    • Timarcha tenebricosa bulgarica
                    • Timarcha tenebricosa heyrowskyi Bechyné, 1945
                    • Timarcha tenebricosa moravica Bechyné, 1945
                    • Timarcha tenebricosa motschulskyi Bechyné, 1945
                    • Timarcha tenebricosa normandiana Bechyné, 1945
                    • Timarcha tenebricosa pauperata Bechyné, 1945
                    • Timarcha tenebricosa submontana Bechyné, 1945
                    • Timarcha tenebricosa tenebricosa (Fabricius, 1775)
                    • Timarcha tenebricosa transgrediens Bechyné, 1948
                  • Timarcha tenuicornis Fairmaire, 1880
                  • Timarcha validicornis Fairmaire, 1873
                    • Timarcha validicornis camoensii Fairmaire, 1873
                    • Timarcha validicornis validicornis Fairmaire, 1873
                • Underslekten Timarchostoma Motschulsky, 1860
                  • Timarcha affinis Laboissiere, 1939
                  • Timarcha aragonica Balbi, 1892
                    • Timarcha aragonica aragonica Balbi, 1892
                    • Timarcha aragonica spectacula Bechyné, 1962
                  • Timarcha asturiensis Kraatz, 1879
                    • Timarcha asturiensis asturiensis Kraatz, 1879
                    • Timarcha asturiensis convexifrons Fairmaire, 1880
                  • Timarcha catalaunensis Fairmaire, 1873
                    • Timarcha catalaunensis belchitensis Bechyné, 1948
                    • Timarcha catalaunensis catalaunensis Fairmaire, 1873
                  • Timarcha chloropus (Germar, 1824)
                    • Timarcha chloropus chloropus (Germar, 1824)
                    • Timarcha chloropus semilaevis Fairmaire, 1873
                  • Timarcha cornuta Bechyné, 1944
                  • Timarcha cyanescens Fairmaire, 1861
                    • Timarcha cyanescens aequata Bechyné, 1953
                    • Timarcha cyanescens cyanescens Fairmaire, 1861
                    • Timarcha cyanescens leseleuci Marseul, 1887
                    • Timarcha cyanescens vizcayensis Bechyné, 1953
                  • Timarcha daillei Laboissiere, 1939
                    • Timarcha daillei cylindricollis Bechyné, 1948
                    • Timarcha daillei daillei Laboissiere, 1939
                    • Timarcha daillei franzi Bechyné, 1962
                    • Timarcha daillei pseudoperezi Bechyné, 1953
                    • Timarcha daillei pygmaeola Bechyné, 1953
                    • Timarcha daillei rhilensis Weise, 1891
                  • Timarcha durmitoriensis Apfelbeck, 1912
                  • Timarcha elliptica Fairmaire, 1873
                  • Timarcha fracassii Meier, 1900
                  • Timarcha geniculata (Germar, 1824)
                  • Timarcha globipennis Fairmaire, 1873
                  • Timarcha goettingensis (Linnaeus, 1758)
                    • Timarcha goettingensis aerea Herrich-Schäffer, 1838
                    • Timarcha goettingensis andrzejowskii Krynicki, 1832
                    • Timarcha goettingensis arvernensis Laboissiere, 1939
                    • Timarcha goettingensis atramentaria Bechyné, 1948
                    • Timarcha goettingensis carpathica Bechyné, 1946
                    • Timarcha goettingensis gallica Fairmaire, 1873
                    • Timarcha goettingensis goettingensis (Linnaeus, 1758)
                    • Timarcha goettingensis mangaliana Bechyné, 1946
                    • Timarcha goettingensis normanna Reiche, 1872
                    • Timarcha goettingensis obenbergeri Bechyné, 1947
                    • Timarcha goettingensis semirufa Pic, 1931
                    • Timarcha goettingensis splendiflora Bechyné, 1946
                  • Timarcha gougeleti Fairmaire, 1859
                  • Timarcha interstitialis Fairmaire, 1861
                    • Timarcha interstitialis bruleriei Bellier, 1870
                    • Timarcha interstitialis cebennica Jeanne, 1965
                    • Timarcha interstitialis interstitialis Fairmaire, 1861
                    • Timarcha interstitialis sinuatocollis Fairmaire, 1861
                  • Timarcha janthinipes Fairmaire, 1880
                  • Timarcha lusitanica (Fabricius, 1781)
                  • Timarcha maritima Perris, 1850
                  • Timarcha melitensis Weise, 1882
                  • Timarcha monserratensis Bechyné, 1962
                  • Timarcha montana Fairmaire, 1873
                  • Timarcha monticola Dufour, 1843
                    • Timarcha monticola altimontana Bechyné, 1948
                    • Timarcha monticola monticola Dufour, 1843
                  • Timarcha oblongula Fairmaire, 1880
                  • Timarcha obsoleta Laboissiere, 1937
                  • Timarcha olivieri Fairmaire, 1868
                    • Timarcha olivieri dubitabilis Fairmaire, 1873
                    • Timarcha olivieri olivieri Fairmaire, 1868
                    • Timarcha olivieri parnassia Fairmaire, 1868
                    • Timarcha olivieri paulmeyeri Stiller, 1926
                  • Timarcha perezi Fairmaire, 1884
                    • Timarcha perezi perezi Fairmaire, 1884
                    • Timarcha perezi praecox Bechyné, 1953
                    • Timarcha perezi vivesi Bechyné, 1962
                  • Timarcha pontavicei Marseul, 1883
                  • Timarcha pratensis Duftschmid, 1825
                  • Timarcha recticollis Fairmaire, 1891
                  • Timarcha sardea Villa, 1835
                  • Timarcha sicelides Reiche, 1860
                  • Timarcha sphaeroptera Fairmaire, 1873
                  • Timarcha stepaneki Bechyné, 1946
                  • Timarcha strangulata Fairmaire, 1861
                    • Timarcha strangulata aubryi Jeanne, 1965
                    • Timarcha strangulata breiti Bechyné, 1953
                    • Timarcha strangulata decens Bechyné, 1953
                    • Timarcha strangulata epipleuralis Bechyné, 1953
                    • Timarcha strangulata pseudostrangulata Bechyné, 1953
                    • Timarcha strangulata sculptipennisBechyné, 1948
                    • Timarcha strangulata strangulata Fairmaire, 1861
                  • Timarcha temperei Jeanne, 1965
                  • Timarcha trapezicollis Fairmaire, 1873
                • Timarcha achardi Bechyné, 1945
                • Timarcha aguilari Bechyné, 1946
                • Timarcha amethystipes Chevrolat, 1873
                • Timarcha angusta Bechyné, 1944
                  • Timarcha angusta algirica Bechyné
                  • Timarcha angusta angusta Bechyné, 1944
                • Timarcha apicicornis Bechyné, 1948
                • Timarcha araneiformis Bechyné, 1947
                • Timarcha atlantica Bechyné, 1944
                • Timarcha brachydera Fairmaire, 1873
                • Timarcha brevipennis Pic, 1919
                • Timarcha corallipes Fairmaire, 1873
                • Timarcha corpulenta Bechyné, 1944
                • Timarcha crassaticollis Fairmaire, 1873
                • Timarcha cyrenaica Peyerimhoff, 1925
                • Timarcha djelfaica Bechyné, 1947
                • Timarcha endora Buquet, 1840
                  • Timarcha endora demoflysi Normand, 1950
                  • Timarcha endora endora Buquet, 1840
                • Timarcha fossulata Bechyné, 1948
                • Timarcha freyi Bechyné, 1953
                • Timarcha generosa Erichson, 1841
                  • Timarcha generosa generosa Erichson, 1841
                  • Timarcha generosa mesatlantica Kocher
                  • Timarcha generosa rungsi Kocher, 1959
                  • Timarcha generosa subscabra Kocher, 1959
                • Timarcha globulata Fairmaire, 1874
                • Timarcha gnaptoroides Bechyné, 1944
                • Timarcha henonii Fairmaire, 1873
                • Timarcha iblanensis Antoine, 1939
                • Timarcha immarginata
                • Timarcha insignis Guerin, 1859
                  • Timarcha insignis inornata Bechyné
                  • Timarcha insignis insignis Guerin, 1859
                • Timarcha khalouensis Jolivet, 1966
                • Timarcha laevigata (Linnaeus, 1767)
                  • Timarcha laevigata inaequalis Pic, 1924
                  • Timarcha laevigata laevigata (Linnaeus, 1767)
                  • Timarcha laevigata robustior Bechyné, 1944
                • Timarcha latipes (Linnaeus, 1767)
                  • Timarcha latipes djerbensis Pic
                  • Timarcha latipes latipes (Linnaeus, 1767)
                • Timarcha mairei Peyerimhoff, 1923
                  • Timarcha mairei humberti Peyerimhoff, 1925
                  • Timarcha mairei lecerfi Kocher, 1951
                  • Timarcha mairei mairei Peyerimhoff, 1923
                  • Timarcha mairei peyerimhoffi Kocher, 1951
                  • Timarcha mairei tichuktensis Kocher, 1957
                • Timarcha marani Bechyné, 1948
                • Timarcha maroccana Weise, 1882
                • Timarcha melas Bechyné, 1944
                • Timarcha nitidipes Bechyné, 1944
                • Timarcha opacicollis Bechyné, 1944
                • Timarcha peresa Peyerimhoff, 1908
                • Timarcha pfefferi Bechyné, 1945
                • Timarcha prujai Kocher, 1963
                • Timarcha punctella Marseul, 1870
                  • Timarcha punctella antiatlantica Jolivet, 1968
                  • Timarcha punctella punctella Marseul, 1870
                  • Timarcha punctella sarhroensis Jolivet, 1968
                  • Timarcha punctella schrammi Kocher, 1956
                  • Timarcha punctella teluetica Escalera, 1914
                • Timarcha punica Lucas, 1849
                • Timarcha reticulata
                • Timarcha riffensis Fairmaire, 1887
                • Timarcha rugosa (Linnaeus, 1767)
                  • Timarcha rugosa aterrima Bechyné, 1944
                  • Timarcha rugosa rugata Kocher, 1951
                  • Timarcha rugosa rugosa (Linnaeus, 1767)
                  • Timarcha rugosa kourili Bechyné, 1944
                  • Timarcha rugosa minutissima Bechyné
                • Timarcha rugosoidea Jolivet, 1944
                • Timarcha scabra (Olivier, 1807)
                • Timarcha scabripennis Fairmaire, 1868
                  • Timarcha scabripennis bechynei Kocher, 1959
                  • Timarcha scabripennis ifranensis Jolivet, 1968
                  • Timarcha scabripennis infoveolata Kocher, 1951
                  • Timarcha scabripennis scabripennis Fairmaire, 1868
                  • Timarcha scabripennis vazquezi Jolivet, 1968
                • Timarcha scutellaris
                • Timarcha sphaeromorpha Bechyné
                • Timarcha spuria Bechyné, 1947
                • Timarcha staneki Bechyné, 1944
                • Timarcha taborskyi Bechyné, 1944
                • Timarcha tangeriana Bechyné, 1943
                • Timarcha tarsata Bechyné, 1944
                • Timarcha theryi Kocher, 1958
                • Timarcha tingitana Fairmaire, 1873
                • Timarcha tunisea Pic, 1919
                • Timarcha turbida Erichson, 1841
                • Timarcha variolosa Bechyné, 1944
                • Timarcha ventricosa Weise, 1882
                  • Timarcha ventricosa aurulenta Kocher, 1957
                  • Timarcha ventricosa mima Bechyné, 1947
                  • Timarcha ventricosa ventricosa Weise, 1882
                • Timarcha zavadili Bechyné, 1944

Referanser

  1. ^ a b Gómez-Zurita, J. (2008) Species and speciation in Timarcha. I: P. Jolivet, J. Santiago-Blay og M. Schmitt (red.) Research on Chrysomelidae. Brill Academic Publishers, Leiden, Nederland.[1]

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