Endomychidae

Family of beetles

Endomychidae
Temporal range: Cenomanian–Recent
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Brachytrycherus humeralis
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Cucujiformia
Superfamily: Coccinelloidea
Family: Endomychidae
Leach, 1815
Type genus
Endomychus
Synonyms

Merophysiidae Seidlitz, 1872

Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.[2]

The subfamilies that are included:

Genera

These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):

  • Acinaces Gerstaecker, 1858 g
  • Aclemmysa Reitter, 1904 g
  • Agaricophilus Motschulsky, 1838 g
  • Amphisternus Germar, 1843 g
  • Amphistethus Strohecker, 1964 g
  • Ancylopus Costa, 1854 g
  • Anidrytus Gerstaecker, 1858 i c g
  • Aphorista Gorham, 1873 i c g b
  • Archipines Strohecker, 1953 i c g
  • Asymbius Gorham, 1896 g
  • Atopomychus Tomaszewska & Szawaryn, 2013 g
  • Atrichonota Arrow, 1925 g
  • Avencymon Strohecker, 1971 g
  • Beccariola Arrow, 1943 g
  • Bolbomorphus Gorham, 1888 g
  • Brachytrycherus Arrow, 1920 g
  • Bystodes Strohecker, 1953 g
  • Cholovocera Motschulsky, 1838 g
  • Chondria Gorham, 1888 g
  • Corynomalus Chevrolat in Dejean, 1836 i c g
  • Cyclotoma Mulsant, 1851 i g
  • Danae Reiche, 1847 i c g b
  • Danascelis Tomaszewska, 1999 i c g
  • Dapsa Latreille, 1829 g
  • Dexialia Sasaji, 1970 g
  • Dialexia Gorham, 1891 g
  • Discolomopsis Shockley, 2006 g
  • Displotera Reitter, 1877 g
  • Ectomychus Gorham, 1888 g
  • Eidorus g
  • Endomychus Panzer, 1795 i c g b
  • Ephebus Chevrolat in Dejean, 1836 i c g
  • Epipocus Germar, 1843 i c g b
  • Epopterus Chevrolat in Dejean, 1836 i c g
  • Eucteanus Gerstaecker, 1857 g
  • Eucymon g
  • Eumorphus Weber, 1801 g
  • Geoendomychus Lea, 1922 g
  • Glesirhanis Shockley & Alekseev, 2014 g
  • Hadromychus Bousquet & Leschen, 2002 i c g b
  • Holoparamecus Curtis, 1833 i c g b
  • Hylaia Guerin, 1857 g
  • Hylaperdina Tomaszewska, 2012 g
  • Idiophyes Blackburn, 1896 g
  • Indalmus Gerstaecker, 1858 g
  • Leiestes Chevrolat, 1836 g
  • Lycoperdina Latreille, 1807 i c g b
  • Meilichius Gerstaecker, 1857 g
  • Merophysia Lucas, 1852 g
  • Microxenus Wallaston, 1861 g
  • Mycetina Mulsant, 1846 i c g b
  • Mychothenus Strohecker, 1953 g
  • Natalinus Tomaszewska, 2011 g
  • Ohtaius Chûjô, 1938 g
  • Palaeoestes Kirejtshuk & Nel, 2009 g
  • Paniegena Heller, 1916 g
  • Parasymbius Arrow, 1920 g
  • Phymaphora Newman, 1838 i c g b
  • Phymaphoroides Motschoulsky, 1857 g
  • Pleganophorus Hampe, 1855 g
  • Polymus Mulsant, 1846 g
  • Pseudindalmus Arrow, 1920 g
  • Rhanidea Strohecker, 1953 i c g b
  • Saula Gerstaecker, 1858 i c g
  • Sinocymbachus Strohecker & Chujo, 1970 g
  • Stenotarsus Perty, 1832 i c g b
  • Stethorhanis Blaisdell, 1931 i c g
  • Trochoideus Westwood, 1833 i c g b
  • Trycherus Gerstaecker, 1857 g
  • Xenomycetes Horn, 1880 i c g b

Data sources: i = ITIS,[3] c = Catalogue of Life,[4] g = GBIF,[5] b = Bugguide.net[6]

Fossil genera

  • Burmalestes Tomaszewska and Ślipiński, 2018[7] Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
  • Cretolestes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
  • Cretaparamecus Tomaszewska, Ślipiński, Bai and Zhang, 2018[7] Burmese amber, Myanmar, Cenomanian
  • Palaeomycetes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
  • Zemyna Tomaszewska, 2018[7] (replaced name Laima Alekseev and Tomaszewska, 2018): Baltic amber, Eocene

References

  1. ^ Tomaszewska, K.Wioletta (2000). "Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)" (PDF). Annales Zoologici. 50 (4): 449–558.
  2. ^ Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B; Whiting, Michael F; McHugh, Joseph V (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)" (PDF). Systematic Entomology. 40 (4): 745. doi:10.1111/syen.12138. S2CID 55206626.
  3. ^ "Endomychidae Report". Integrated Taxonomic Information System. Retrieved 2018-05-07.
  4. ^ "Browse Endomychidae". Catalogue of Life. Archived from the original on 2021-06-18. Retrieved 2018-05-07.
  5. ^ "Endomychidae". GBIF. Retrieved 2018-05-07.
  6. ^ "Endomychidae Family Information". BugGuide.net. Retrieved 2018-05-07.
  7. ^ a b c d e Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018). "The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber". Cretaceous Research. 91: 287–298. doi:10.1016/j.cretres.2018.07.001. S2CID 135401188.

External links

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Extant Coleoptera families
Suborder Archostemata
  • Crowsoniellidae (Crowsoniella relicta)
  • Cupedidae (reticulated beetles)
  • Jurodidae (Sikhotealinia zhiltzovae)
  • Micromalthidae (telephone-pole beetle)
  • Ommatidae
Suborder Adephaga
Extant families
  • Amphizoidae (trout-stream beetles)
  • Aspidytidae
  • Carabidae (ground beetles)
  • Cicindelidae (tiger beetles)
  • Dytiscidae (predaceous diving beetles)
  • Gyrinidae (whirligig beetles)
  • Haliplidae (crawling water beetles)
  • Hygrobiidae
  • Meruidae (Meru phyllisae)
  • Noteridae (burrowing water beetles)
  • Trachypachidae (false ground beetles)
Suborder Myxophaga
Suborder Polyphaga
Bostrichiformia
Bostrichoidea
  • Bostrichidae (auger beetles)
  • Dermestidae (skin beetles)
  • Endecatomidae
  • Jacobsoniidae (Jacobson's beetles)
  • Nosodendridae (wounded-tree beetles)
  • Ptiniidae (furniture beetles, death watch beetles, spider beetles)
Derodontoidea
  • Derodontidae (tooth-necked fungus beetles)
Cucujiformia
Chrysomeloidea
Cleroidea
Coccinelloidea
Cucujoidea
Curculionoidea
(weevils)
  • Anthribidae (fungus weevils)
  • Attelabidae (leaf-rolling weevils)
  • Belidae (primitive weevils)
  • Brentidae (straight snout weevils, New York weevil)
  • Caridae
  • Curculionidae (true weevils, bark beetles, ambrosia beetles)
  • Nemonychidae (pine flower weevils)
Lymexyloidea
  • Lymexylidae (ship-timber beetles)
Tenebrionoidea
  • Aderidae (ant-like leaf beetles)
  • Anthicidae (ant-like flower beetles)
  • Archeocrypticidae (cryptic fungus beetles)
  • Boridae (conifer bark beetles)
  • Chalcodryidae
  • Ciidae (minute tree-fungus beetles)
  • Melandryidae (false darkling beetles)
  • Meloidae (blister beetles)
  • Mordellidae (tumbling flower beetles)
  • Mycetophagidae (hairy fungus beetles)
  • Mycteridae (palm and flower beetles)
  • Oedemeridae (false blister beetle)
  • Perimylopidae, or Promecheilidae
  • Prostomidae (jugular-horned beetles)
  • Pterogeniidae
  • Pyrochroidae (fire-coloured beetles)
  • Pythidae (dead log bark beetles)
  • Ripiphoridae (wedge-shaped beetles)
  • Salpingidae (narrow-waisted bark beetles)
  • Scraptiidae (false flower beetles)
  • Stenotrachelidae (false longhorn beetles)
  • Synchroidae (synchroa bark beetles)
  • Tenebrionidae (darkling beetles)
  • Tetratomidae (polypore fungus beetles)
  • Trictenotomidae
  • Ulodidae
  • Zopheridae (ironclad beetles, cylindrical bark beetles)
Elateriformia
Buprestoidea
  • Buprestidae (jewel beetles, or metallic wood-boring beetles)
  • Schizopodidae
Byrrhoidea
  • Byrrhidae (pill beetles)
  • Callirhipidae (cedar beetles)
  • Chelonariidae (turtle beetles)
  • Cneoglossidae
  • Dryopidae (long-toed water beetles)
  • Elmidae (riffle beetles)
  • Eulichadidae (forest stream beetles)
  • Heteroceridae (variegated mud-loving beetles)
  • Limnichidae (minute mud beetles)
  • Lutrochidae (travertine beetles)
  • Psephenidae (water-penny beetles)
  • Ptilodactylidae
Dascilloidea
  • Dascillidae (soft bodied plant beetles)
  • Rhipiceridae (cicada beetle, cicada parasite beetles)
Elateroidea
  • Artematopodidae (soft-bodied plant beetles)
  • Brachypsectridae (Texas beetles)
  • Cantharidae (soldier beetles)
  • Cerophytidae (rare click beetles)
  • Elateridae (click beetles)
  • Eucnemidae (false click beetles)
  • Jurasaidae
  • Lampyridae (fireflies)
  • Lycidae (net-winged beetles)
  • Omethidae (false fireflies, long-lipped beetles)
  • Phengodidae (glowworm beetles)
  • Rhagophthalmidae
  • Sinopyrophoridae
  • Throscidae (false metallic wood-boring beetles)
Rhinorhipoidea
  • Rhinorhipidae (Rhinorhipus tamborinensis)
Scirtoidea
Scarabaeiformia
Scarabaeoidea
  • Belohinidae (Belohina inexpectata)
  • Bolboceratidae
  • Diphyllostomatidae (false stag beetles)
  • Geotrupidae (dor beetles)
  • Glaphyridae (bumble bee scarab beetles)
  • Glaresidae (enigmatic scarab beetles)
  • Hybosoridae (scavenger scarab beetles)
  • Lucanidae (stag beetles)
  • Ochodaeidae (sand-loving scarab beetles)
  • Passalidae (betsy beetles)
  • Pleocomidae (rain beetles)
  • Scarabaeidae (scarabs)
  • Trogidae (hide beetles)
Staphyliniformia
Histeroidea
  • Histeridae (clown beetles)
  • Sphaeritidae (false clown beetles)
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Hydrophiloidea
Staphylinoidea
  • Agyrtidae (primitive carrion beetles)
  • Hydraenidae
  • Leiodidae (round fungus beetles)
  • Ptiliidae (feather-winged beetles)
  • Silphidae (carrion beetles)
  • Staphylinidae (rove beetles)
Taxon identifiers
Endomychidae
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