Ulmus 'Toledo'

Elm cultivar

Ulmus 'Toledo'
'Toledo'
Genus	Ulmus
Cultivar	'Toledo'
Origin	Spain

The elm cultivar Ulmus 'Toledo' was raised from seed collected in 1999 from a tree believed to be Ulmus minor growing in the eponymous city (39°51′51″N 4°1′30″W / 39.86417°N 4.02500°W / 39.86417; -4.02500),^[1] by researchers at the Escuela Técnica Superior de Ingenieros de Montes, Universidad Politėcnica de Madrid. However, retrospective DNA analysis revealed that the clone also had Ulmus pumila DNA, the species introduced to Spain at the end of the 16th century,^[1] and has consequently been dropped from the research programme.^[2]

'Toledo' was introduced to the UK in 2020 as part of an assessment by Butterfly Conservation of DED-resistant cultivars as potential hosts of the endangered White-letter Hairstreak.^[2]

Description

'Toledo' grew at a comparatively fast rate of 89 cm per annum in the trials at Puerta de Hierro, Madrid. The erect branches form an irregular crown and are devoid of corky tissue. The leaves, on 6 mm petioles, are ovate, typically acuminate at the apex, the average length and width 47 × 27 mm, the margins doubly serrate. Foliar density relative to 'Sapporo Autumn Gold' is described as 'high'. In the Madrid study, the appearance of the tree was rated 2.9 / 5. ^[1] The leaves flush in early April (week 14) in southern England.

Pests & diseases

'Toledo' was found to have a very high resistance to Dutch Elm Disease, on a par with, if not greater than, the hybrid cultivar 'Sapporo Autumn Gold'.^[1]

Accessions

Europe

Grange Farm Arboretum, Lincolnshire, UK. Acc. no. not known.
Great Fontley Farm, Hampshire, UK. Butterfly Conservation elm trial plantation. One small sapling planted 2020.

References

^ ^a ^b ^c ^d Martín, JA; Solla, A; Venturas, M; Collada, C; Domínguez, J; Miranda, E; Fuentes, P; Burón, M; Iglesias, S; Gil, L (2015-04-01). "Seven Ulmus minor clones tolerant to Ophiostoma novo-ulmi registered as forest reproductive material in Spain". IForest - Biogeosciences and Forestry. 8 (2). Italian Society of Sivilculture and Forest Ecology (SISEF): 172–180. doi:10.3832/ifor1224-008. ISSN 1971-7458.
^ ^a ^b Brookes, A. H. (2020). Great Fontley Elm Trial, 2020 Report. Butterfly Conservation, Lulworth, UK.

Elm species, varieties, hybrids, hybrid cultivars and species cultivars

Species, varieties and subspecies

U. alata (Winged elm)
U. americana (American elm)
U. americana var. floridana (Florida elm)
U. bergmanniana (Bergmann's elm)
U. bergmanniana var. bergmanniana
U. bergmanniana var. lasiophylla
U. castaneifolia (Chestnut-leafed or multinerved elm)
U. changii (Hangzhou elm)
U. changii var. changii
U. changii var. kunmingensis (Kunming elm)
U. chenmoui (Chenmou or Langya Mountain elm)
U. chumlia
U. crassifolia (Cedar or Texas cedar elm)
U. davidiana (David or Father David elm)
U. davidiana var. davidiana
U. davidiana var. japonica (Japanese elm)
U. elongata (Long raceme elm)
U. gaussenii (Anhui or hairy elm)
U. glabra (Wych or scots elm)
U. glaucescens (Gansu elm)
U. glaucescens var. glaucescens
U. glaucescens var. lasiocarpa (hairy-fruited glaucescent elm)
U. harbinensis (Harbin elm)
U. ismaelis
U. laciniata (Manchurian cut-leaf or lobed elm)
U. laciniata var. nikkoensis (Nikko elm)
U. laevis (European white elm)
U. laevis var. celtidea
U. laevis var. parvifolia
U. laevis var. simplicidens
U. lamellosa (Hebei elm)
U. lanceifolia (Vietnam elm)
U. macrocarpa (Large-fruited elm)
U. macrocarpa var. glabra
U. macrocarpa var. macrocarpa
U. mexicana (Mexican elm)
U. microcarpa (Tibetan elm)
U. minor (Field elm)
U. minor subsp. minor
U. minor var. italica
U. parvifolia (Chinese or lacebark elm)
U. parvifolia var. coreana (Korean elm)
U. prunifolia (Cherry-leafed elm)
U. pseudopropinqua (Harbin spring elm)
U. pumila (Siberian elm)
U. rubra (Slippery elm)
U. serotina (September elm)
U. szechuanica (Szechuan (Sichuan) or red-fruited elm)
U. thomasii (Rock or cork elm)
U. uyematsui (Alishan elm)
U. villosa (Cherry-bark or marn elm)
U. wallichiana (Himalayan or kashmir elm)
U. wallichiana subsp. wallichiana
U. wallichiana subsp. xanthoderma
U. wallichiana var. tomentosa

Disputed species, varieties and subspecies

U. boissieri
U. minor subsp. canescens (Grey, grey-leafed or hoary elm)
U. elliptica

Hybrids

U. davidiana var. japonica × U. minor
U. × arbuscula
U. × arkansana
U. × brandisiana
U. × diversifolia
U. × hollandica (Dutch elm)
U. × hollandica var. insularum
U. × intermedia
U. × mesocarpa

Species cultivars

American elm	American Liberty Ascendens Augustine Aurea Beaverlodge Beebe's Weeping Brandon Burgoyne JFS Prince II College Columnaris Creole Queen Deadfree Delaware Exhibition Fiorei Flick's Spreader Folia Aurea Variegata Great Plains Hines Incisa Independence Iowa State Jackson Jefferson Kimley Klehmii L'Assomption Lake City Lewis & Clark Littleford Maine Markham Miller Park Minneapolis Park Moline Morden New Harmony Nigricans Patmore Pendula Penn Treaty Princeton Pyramidata Queen City Sheyenne Skinner Upright St. Croix Star Valley Forge Variegata Vase Washington
Cedar elm	Brazos Rim
Chinese elm	A-1 A. Ross Central Park Blizzard BSNUPF Burgundy Burnley Select Catlin Chessins Churchyard Cork Bark D. B. Cole Drake Dynasty Ed Wood Elsmo Emer I Emer II Emerald Prairie Frosty Garden City Clone Geisha Glory Golden Rey Hallelujah Hokkaido Jade Empress King's Choice Littleleaf Lois Hole Matthew Milliken Nire-keyaki Ohio Orange Ribbon Pathfinder Pendens Prairie Shade Prince Richard Red Fall Sabamiki Sagei Seiju Select 380 Sempervirens Small Frye State Fair Stone's Dwarf Taiwan The Thinker Todd True Green UPMTF Ware's Yarralumla Yatsubusa Zettler Ulmus parvifolia f. lanceolata
European white elm	Aureovariegata Colorans Helena Ornata Pendula Punctata Urticifolia
Field elm	Ademuz Albo-Dentata Amplifolia Argenteo-Variegata Atinia Atinia Pyramidalis Atinia Variegata Bea Schwarz Biltii Christine Buisman Concavaefolia Coritana Cretensis Cucullata Cucullata Variegata Dehesa de Amaniel Dehesa de la Villa Dicksonii Dijkwel Erecta Folia Alba-Punctata Glandulosa Goodyeri Hoersholmiensis Holmstruph Hunnybunii Hunnybunii pseudo-Stricta Laciniata Lanuginosa Majadahonda Microphylla Pendula Microphylla Purpurea Microphylla Rubra Monumentalis Pendula Picturata Plotii Propendens Punctata Purpurascens Purpurea Retiro Reverti Rugosa Rueppellii Sarniensis Schuurhoek Silvery Gem Sowerbyi Stricta Suberosa Umbraculifera Gracilis Umbraculifera Viminalis Viminalis Aurea Viminalis Betulaefolia Viminalis Gracilis Viminalis Incisa Viminalis Marginata Viminalis Pendula Viminalis Pulverulenta Viminalis Stricta Virgata Webbiana
Japanese elm	Discovery Freedom Jacan JFS-Bieberich Mitsui Centennial Prospector Reperta Reseda Thomson Validation
Siberian elm	Ansaloni Aurea Aurescens Chinkota Dropmore Dwarf Weeper Green King Hansen Harbin Manchu Mauro Mr. Buzz Park Royal Pendula Pinnato-ramosa Poort Bulten Puszta Pyramidalis Fiorei Zhonghua Jinye
Winged elm	Lace Parasol
Wych elm	Albo-Variegata Australis Camperdownii Cebennensis Concavaefolia Cornuta Corylifolia Purpurea Corylifolia Dovaei Fastigiata Macrophylla Fastigiata Stricta Fastigiata Variegata Firma Flava Gigantea Gittisham Grandidentata Holgeri Horizontalis Insularis Latifolia Latifolia Aurea Latifolia Aureo-Variegata Latifolia Nigricans Luteo Variegata Lutescens Macrophylla Maculata Minor Nana Nigra Nitida Oblongata Pendula Macrophylla Pendula Variegata Pyrenaica Spectabilis Superba Tomentosa

Hybrid cultivars

Androssowii Amsterdam Arno Cathedral Clusius Columella Den Haag Dodoens Escaillard Fiorente Frontier Fuente Umbria Homestead Karagatch Lobel Morfeo Morton Morton Glossy Morton Plainsman Morton Red Tip Morton Stalwart Nanguen New Horizon Patriot Plantyn Plinio Rebella Rebona Recerta Regal Repura Revera San Zanobi Sapporo Autumn Gold Sapporo Gold 2 Stavast Toledo Urban Wanoux Wingham
Dutch elm	Alba Angustifolia Balder Belgica Blandford Canadian Giant Cicestria Cinerea Commelin Dampieri Dauvessei Daveyi Dumont Elegantissima Eleganto-Variegata Etrusca Fastigiata Fjerrestad Folia Rhomboidea Freja Fulva Gaujardii Groeneveld Haarlemensis Klemmer Loke Macrophylla Aurea Major Microphylla Modiolina Muscaviensis Odin Pioneer Pitteurs Serpentina Smithii Superba Tricolor Tyr Vegeta Viminalis Viscosa Wentworthii Pendula Wredei Ypreau
U. × intermedia	Coolshade Fremont Improved Coolshade Lincoln Rosehill Willis